Endangered and Threatened Wildlife; Notice of 12-Month Finding on a Petition To List the Sperm Whale (Physeter macrocephalus, 68032-68037 [2013-27180]
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Federal Register / Vol. 78, No. 219 / Wednesday, November 13, 2013 / Notices
public comment received.5 Most
recently, on May 9, 2013, the President
announced competitions to create three
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institutes, and the Administration
continues to call on Congress to act on
the President’s proposal and FY 2014
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Request for Comments: The AMNPO
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National Network for Manufacturing
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These documents address topics
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Documents related to additional high
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See the FOR FURTHER INFORMATION
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Dated: November 6, 2013.
Phillip Singerman,
Associate Director for Innovation and
Industry Services.
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2013/05/09/obama-administration-launchescompetition-three-new-manufacturing-innova.
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DEPARTMENT OF COMMERCE
National Oceanic and Atmospheric
Administration
[Docket No. 1206013325–3912–03]
RIN 0648–XA983
Endangered and Threatened Wildlife;
Notice of 12-Month Finding on a
Petition To List the Sperm Whale
(Physeter macrocephalus) as an
Endangered or Threatened Distinct
Population Segment (DPS) in the Gulf
of Mexico
National Marine Fisheries
Service (NMFS), National Oceanic and
Atmospheric Administration (NOAA),
Commerce.
ACTION: Status review; notice of finding.
AGENCY:
We, NMFS, announce a 12month finding on a petition to list the
sperm whale (Physeter macrocephalus)
in the Gulf of Mexico as an endangered
or threatened distinct population
segment (DPS) under the Endangered
Species Act of 1973 as amended (ESA).
We conducted a review of the status of
this population, as described below.
Based on the best available scientific
and commercial information, we find
that the petitioned action is not
warranted.
DATES: The finding announced in this
notice was made on November 13, 2013.
ADDRESSES: Information used to make
this finding is available for public
inspection by appointment during
normal business hours at NMFS
Headquarters, Protected Resources
Office, 1315 East-West Highway, Silver
Spring, MD 20910. This file includes the
information provided by the public and
scientific and commercial information
gathered for the status review. The
petition and a list of the references we
used can also be found at https://
www.nmfs.noaa.gov/pr/.htm.
FOR FURTHER INFORMATION CONTACT:
Marta Nammack, NMFS, Office of
Protected Resources, (301) 427–8469.
SUPPLEMENTARY INFORMATION: On
December 9, 2011, we received a
petition from WildEarth Guardians to
list the sperm whale (Physeter
macrocephalus) population in the Gulf
of Mexico as an endangered or
threatened Distinct Population Segment
(DPS) under the Endangered Species
Act (ESA); sperm whales are currently
listed as a single endangered species
throughout their global range (35 FR
8495; June 2, 1970). The petitioner also
requested designation of critical habitat
concurrent with the listing.
After reviewing the petition, the
literature cited in the petition, and other
SUMMARY:
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literature and information available in
our files, we found that the petition met
the requirements of the regulations
under 50 CFR 424.14(b)(2) and
determined that the petition presented
substantial information indicating that
the petitioned action may be warranted
(78 FR 19176; March 29, 2013). At that
time, we commenced a status review of
the sperm whale in the Gulf of Mexico
and solicited information pertaining to
the population. Section 4(b)(3)(B) of the
ESA requires that when a petition to
revise the List of Endangered and
Threatened Wildlife and Plants is found
to present substantial scientific and
commercial information, we make a
finding on whether the petitioned action
is (a) not warranted, (b) warranted, or (c)
warranted but precluded from listing by
other pending proposals of higher
priority. This finding is to be made
within 12 months of the date the
petition was received, and the finding is
to be published promptly in the Federal
Register.
There are two key tasks associated
with conducting an ESA status review.
The first is to determine whether the
petitioned entity qualifies as one or
more species under the ESA. The ESA
defines the term ‘‘species’’ to include
‘‘any distinct population segment of any
species of vertebrate fish or wildlife
which interbreeds when mature.’’ If the
petitioned entity qualifies as a species,
the second task is to conduct an
extinction risk assessment to determine
whether the species is threatened or
endangered. The ESA defines the term
‘‘endangered species’’ as ‘‘any species
which is in danger of extinction
throughout all or a significant portion of
its range.’’ The term ‘‘threatened
species’’ is defined as ‘‘any species
which is likely to become endangered
within the foreseeable future throughout
all or a significant portion of its range.’’
Thus, we interpret an ‘‘endangered
species’’ to be one that is presently in
danger of extinction. A ‘‘threatened
species,’’ on the other hand, is not
presently in danger of extinction, but is
likely to become so in the foreseeable
future (that is, at a later time). In other
words, the primary statutory difference
between a threatened and endangered
species is the timing of when a species
may be in danger of extinction, either
presently (endangered) or in the
foreseeable future (threatened).
Species Background
The sperm whale (Linnaeus, 1758) is
listed as an endangered species under
the ESA. It was first listed under the
precursor to the ESA, the Endangered
Species Conservation Act of 1969, and
remained on the list of threatened and
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endangered species after the passage of
the ESA in 1973 (35 FR 18319;
December 2, 1970). Whaling was the
main reason for listing the sperm whale.
Commercial whaling for this species
ended in 1988 with the implementation
of a moratorium against whaling by the
International Whaling Commission
(IWC). While whaling was eliminated by
the IWC whaling moratorium, several
potential threats remain, as discussed in
the sperm whale recovery plan (NMFS,
2010a). Sperm whales are deep and
prolonged divers and use the entire
water column, even in very deep areas.
Most sperm whales are found in very
deep waters (>3,000 m), but they
generally feed between 500–1,000 m
where most of their prey is found.
Sperm whales feed primarily on largeand medium-sized squid, but the list of
documented food items is fairly long
and diverse, including other
cephalopods and medium-and largesized demersal fish, such as rays,
sharks, and many teleosts (Berzin, 1972;
Clarke 1977, 1980; Rice, 1989). The diet
of large males in some areas, especially
in high northern latitudes, is dominated
by fish (Rice, 1989). Lockyer (1981)
estimated sperm whales consumed
about 3.0–3.5 percent of their body
weight per day.
Sperm whales are perhaps the most
widely distributed mammal species on
Earth. The social organization of most
mammals is characterized by female
philopatry and male dispersal. Groups
of females and juveniles are found
mainly at low latitudes, while males
reach polar waters, returning to tropical
and subtropical waters to breed. Sperm
whales are organized in groups in which
females (some related to each other and
some not) travel with their sub-adult
offspring. Mature female and immature
sperm whales of both sexes are found in
more temperate and tropical waters
from the equator to around 45ßN
throughout the year. Adult males will
move extensively, even to polar waters,
and then return to tropical and
subtropical waters.
Sperm whales mature slowly and can
live to ages in excess of 60 years (Rice,
1989). Females usually begin ovulating
at 7–13 years of age and usually
conceive at about age 9 (Rice, 1989).
Maturation in males usually begins in
this same age interval, but most
individuals do not become fully mature
until their twenties. In the North
Atlantic Ocean, the peak breeding
season for sperm whales occurs during
the spring (March/April to June),
although some mating activity occurs
December to August. In the South
Atlantic the peak breeding season is
presumed to occur in the austral spring.
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During mating seasons, prime bulls in
their late twenties and older rove among
groups of females. Because females
within a group often come into estrus
synchronously, the males need not
remain with the females for the breeding
season to achieve maximal breeding
success (Best and Butterworth, 1980)
and their association with a group can
be as brief as several hours. Gestation
lasts well over a year, with credible
estimates of the normal duration ranging
from 15 months to more than a year and
a half. Lactation lasts at least 2 years,
and the inter-birth-interval is 4–6 years
(Best et al., 1984) for prime-aged
females. Female sperm whales rarely
become pregnant after the age of 40
(Whitehead, 2003). Two particular
aspects of the sperm whale’s
reproductive biology are relevant to
recovery. First, the maximal rate of
increase in reproduction is very low,
perhaps no more than one or two
percent per year. Second, selective
killing of large males by modern
whaling could have had the residual
effect of reducing reproductive rates
(Whitehead et al., 1997).
Status Review
Our 90-day finding accepting the
petition solicited information from the
public and initiated a status review of
the sperm whale in the Gulf of Mexico
(GOM) to gather any additional
information to inform our review of the
petitioned action and our application of
the DPS policy. We reviewed the best
available information, and we
conducted a DPS analysis to determine
whether the GOM population of the
sperm whale qualifies as a DPS under
the ESA. Here we review the best
available information on physical,
physiological, ecological, and
behavioral factors to determine whether
the GOM population is discrete.
Are sperm whales in the Gulf of Mexico
discrete from other sperm whale
populations?
The ESA provides for listing species,
subspecies, or DPSs of vertebrate
species. When we evaluate a petition to
list an entity as threatened or
endangered under the ESA, we must
first determine whether the petitioned
entity qualifies as a species under the
ESA. This petition argues that the Gulf
of Mexico sperm whale population
meets the requirements for being
identified as a DPS and requests we list
sperm whales in the Gulf of Mexico as
a threatened or endangered DPS.
Our joint NMFS–U.S. Fish and
Wildlife Service (USFWS) Policy on
Recognition of Distinct Vertebrate
Population Segments under the
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Endangered Species Act (DPS policy)
(61 FR 4722; February 7, 1996)
identifies two elements that must be
considered when identifying a DPS: (1)
The discreteness of the population
segment in relation to the remainder of
the species (or subspecies) to which it
belongs; and (2) the significance of the
population segment to the species to
which it belongs. A population segment
of a vertebrate species may be
considered discrete if it satisfies either
one of the following conditions: (1) It is
markedly separated from other
populations of the same taxon as a
consequence of physical, physiological,
ecological, or behavioral factors.
Quantitative measures of genetic or
morphological discontinuity may
provide evidence of this separation; or
(2) it is delimited by international
governmental boundaries within which
differences in control of exploitation,
management of habitat, conservation
status, or regulatory mechanisms exist
that are significant in light of section
4(a)(1)(D) of the ESA. If a population
segment is considered discrete by one or
more of the above conditions, its
biological and ecological significance
will then be considered in light of
Congressional guidance (see Senate
Report 151, 96th Congress, 1st Session)
that the authority to list DPSs be used
‘‘. . . sparingly’’ while encouraging the
conservation of genetic diversity. The
DPS policy directs us to consider
available scientific evidence of the
discrete population segment’s
importance to the taxon to which it
belongs. This consideration may
include, but is not limited to, the
following: (1) Persistence of the discrete
population segment in an ecological
setting unusual or unique for the taxon;
(2) evidence that loss of the discrete
population segment would result in a
significant gap in the range of a taxon;
(3) evidence that the discrete population
segment represents the only surviving
natural occurrence of a taxon that may
be more abundant elsewhere as an
introduced population outside its
historic range; or (4) evidence that the
discrete population segment differs
markedly from other populations of the
species in its genetic characteristics.
DPS Analysis
To determine if the sperm whale in
the GOM meets the DPS criteria, we
evaluate the best available information
to determine whether sperm whales in
the Gulf of Mexico are markedly
separated as a consequence of physical,
physiological, ecological, or behavioral
factors from other populations of the
sperm whale.
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Genetics—An examination of the best
available genetic information reveals
that, although there is strong mtDNA
evidence of population structuring
indicating differences between the GOM
population and sperm whales in the
northwest Atlantic, this is not coupled
with nDNA evidence that would
indicate that males from the GOM are
genetically different from males in the
northwest Atlantic. Physically mature
male sperm whales typically range over
huge distances on their own (Best, 1979;
Rice, 1989; Whitehead, 1993; Whitehead
and Weilgart, 2000; Teloni et al., 2008).
In contrast to females, males disperse
from their natal units at a mean
estimated age of 6 years, when they
migrate slowly into higher latitudes
prior to attaining sexual maturity at 18–
21 years (Whitehead and Weilgart,
2000). This is reÖected in high
variability and a lack of geographical
structure in nDNA relative to mtDNA
(Lyrholm et al., 1999).
There are statistically significant
patterns of mtDNA differentiation
between oceans (Engelhaupt, 2004;
SWSS, 2008; Engelhaupt et al., 2009;
NMFS, 2010a); however, studies
examining nDNA reveal either no
significant (Lyrholm et al., 1999) or low
(Bond, 1999) degrees of population
structuring between oceans. Engelhaupt
et al. (2009) suggest that the discrepancy
between mtDNA and nDNA
differentiation may reflect sex biased
dispersal, and male mediated gene flow
may connect geographically isolated
regions on an oceanic scale. Their
analysis of nDNA showed no significant
difference between whales sampled in
the GOM and those from other areas of
the North Atlantic, indicating that
mature males move in and out of the
GOM. The results of the Engelhaupt et
al. (2009) study indicate that population
structuring is different for mtDNA
compared with population structuring
for nDNA.
At best, mtDNA evidence suggests
that females are philopatric; however,
mtDNA does not alone describe
population structure. Because mtDNA is
maternally inherited, differences in
mtDNA haplotypes between
populations do not necessarily mean
that the populations are substantially
reproductively isolated from each other
because they do not provide any
information on males. Due to the wide
ranging nature of mature male sperm
whales, males from one population may
breed with females from other
populations. We have indicated in other
status reviews that mtDNA data may
indicate that populations are discrete,
but in species where female and male
movement patterns differ, nDNA data
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may indicate that the populations are
homogeneous (see e.g., loggerhead sea
turtle, 68 FR 53947, September 15, 2003
at 53950–51 and Conant et al., 2009, at
18, 22, 25–28; southern resident killer
whale, Krahn et al., 2002, at 23–30). As
noted in SWSS (2008), a male sperm
whale tagged in 2002 moved into the
North Atlantic for more than 2 months,
providing the first evidence that the
GOM population may not be a stock
isolated from the North Atlantic (SWSS,
2008; Waring et al., 2012). Its return to
the GOM included an extended stay off
the northwest Cuban coast, and it
summered in two different regions of
the upper GOM and visited the Gulf of
Campeche twice (SWSS, 2008). While
some may view this as support for
separate stocks in the GOM and the
North Atlantic, SWSS (2008) notes that
few males were sampled in the GOM.
Because the tags were deployed from
June to early August, more individuals
were tracked during the summer months
(SWSS, 2008). Therefore, it is likely that
mature males were not in the GOM at
this time, as they spend most of their
time in colder waters at high latitudes
and only visit tropical waters to
reproduce (Best 1979; Whitehead and
Arnbom 1987; Whitehead 2003, as cited
in SWSS (2008)).
The fact that males move in and out
of the GOM and interbreed with females
from other populations when mature, as
evidenced by the homogeneity of the
nDNA, indicates that the GOM
population is not markedly separated
from other populations in the Atlantic
Ocean. Engelhaupt et al. (2009)
demonstrate that a single, undivided
genetic population of sperm whales is
found from the GOM to at least northern
Europe. As we have summarized here,
the best available genetic information
indicates that sperm whales in the GOM
are not discrete from other sperm whale
populations.
Vocalization—We next examined
information on codas. Sperm whale
social structure is complex, with
females, calves, and immature animals
of both sexes living in relatively stable
social ‘‘units’’ containing on average 11–
12 animals that persist for decades
(Rendell and Whitehead, 2004). These
sperm whale social groups
communicate via codas: Repeated
stereotyped sequences of 3–40
broadband (0–16 kHz) clicks generally
heard during periods of socializing
(Watkins and Schevill, 1977). Codas are
shared among individuals of a social
unit and are considered to be primarily
for intra-group communication
(Weilgart and Whitehead, 1997; Rendell
and Whitehead, 2004). These
distinctive, short, patterned series of
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clicks are associated with social
behavior and interactions within social
groups (Weilgart and Whitehead, 1993).
Significant differences in vocalization
or coda repertoire exist amongst smaller
social groups or ‘‘units’’ of sperm
whales, and this variation amongst
social units or groups is commonplace
for sperm whales (Weilgart and
Whitehead, 1997; Rendell and
Whitehead, 2004). Differences in
vocalization are culturally transmitted
by the matrilineal line, and there is a
difference between geographical sperm
whale variation in codas
(macrogeographic) and coda ‘‘dialects’’
(microgeographic) (Mundinger, 1982). In
a study of sperm whales in the southern
Pacific Ocean, Weilgart and Whitehead
(1997) found that the sperm whale
groups they encountered had distinctive
dialects in coda usage based on analyses
of interclick intervals (ICIs), the time
intervals between clicks in a coda,
standardized to total coda length. The
group-specific dialects that are found in
sperm whales have even been deemed
as similar to those which occur in killer
whale ‘‘vocal clans’’ (Weilgart and
Whitehead, 1997; Rendell and
Whitehead, 2003).
Codas and mtDNA have been linked;
a study of six sperm whale groups
revealed a clear link between mtDNA
and coda repertoire as groups with
similar mtDNA tended to have similar
coda usage dialects (Whitehead et al.,
1998). These results indicate codas are
transmitted across generations
matrilineally. Whitehead et al. (1998)
suggested vertical cultural transmission
(offspring learn codas from their
mothers) as the best explanation for this
pattern. This may reflect the mtDNA
information presented above suggesting
population structure, without
consideration of the nDNA. The sperm
whale seismic study (SWSS, 2008) cited
in the petition found variation in
vocalization between the north central
GOM and the northwest GOM. Because
there is evidence of different types of
coda variation (i.e., macrogeographic
versus microgeographic dialects) within
the GOM, communication is passed
down from the mother, and adult male
sperm whales travel outside the Gulf of
Mexico, the communication difference
between GOM sperm whales and sperm
whales from other populations does not
indicate sperm whales in the GOM are
‘‘markedly’’ separate.
Group size—While group size in the
GOM is smaller on average than in other
oceans, group size is variable
throughout their global range. The fact
that group sizes are similar to those in
the Caribbean and smaller than group
sizes in some other oceans (SWSS,
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2008) does not show a ‘‘marked’’
separation from other sperm whale
group sizes. Christal et al. (1998) note
that estimated social unit size in the
Galapagos, for example, ranged from 3
to 24 individuals and presented
evidence of splitting and merging of
units and of transfer of individuals
between units. The considerable
variation in unit size (perhaps caused by
demographic processes) suggests that
the benefits of remaining in a social unit
usually outweigh selection for some
optimal unit size (Christal et al., 1998).
Richter et al. (2008) note that it could
be argued that differences in ecological
conditions in which various sperm
whale populations live are reflected in
the parameters of their social behavior,
such as group size and association rate
(Richter et al., 2008). The best available
evidence does not indicate that sperm
whale group size in the GOM is
different from all other populations of
the sperm whale.
Whale size—Mean size of sperm
whales in the GOM (8.5 m) has been
reported to be smaller than that of other
sperm whale populations (e.g., 10 m for
the Gulf of California population)
(SWSS, 2008). While photographic data
on known males and sound pulse
studies showed that those measured in
the GOM were smaller than breeding
males elsewhere (Jaquet et al., 2006;
Antunes et al., 2006), no mature males
have been observed in the GOM. This
only confirms that younger male whales
that have recently departed from their
mothers are smaller than those at full
maturity, which is not noteworthy.
Older males, which apparently only
pass through the GOM for breeding, are
larger than the younger males that have
not yet migrated out of the GOM.
Further, whale size data from these
studies have never been normalized to
account for age, so a reliable comparison
cannot be made. Finally, Jochens et al.
(2008) argue that female/adolescent size
differences among sperm whale
populations may be the result of nothing
more than differences in prey,
suggesting that ‘‘it is possible that the
population studied is smaller because
smaller animals may prefer the
shallower waters relative to their diving
ability and/or availability of suitable
prey.’’ Whales may assort themselves by
water depths to match their body sizes.
Finally, even if GOM whales are a little
smaller on average than other
populations of sperm whale, such a
modest difference is not sufficient to
demonstrate that the GOM population is
‘‘markedly separated’’ from other sperm
whale populations.
International boundaries—In
examining whether a population is
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discrete based on international
governmental boundaries, we are to
examine differences in the control of
exploitation, management of habitat,
conservation status, or regulatory
mechanisms exist that are significant in
light of section 4(a)(1)(D) of the ESA.
Section 4(a)(1)(D), the inadequacy of
existing regulatory mechanisms, is one
of the five factors we must evaluate to
determine whether to list a species. We
did not find any information pointing to
significant differences in control of
exploitation, management of habitat,
conservation status, or regulatory
mechanisms between the population of
sperm whales in the GOM and any other
particular population of the sperm
whale such that the population of the
sperm whale in the GOM could be
considered discrete from a sperm whale
population outside of the GOM. The
ESA extends prohibitions against take of
endangered species by any person
subject to the jurisdiction of the United
States within the United States, its
territorial waters, or on the high seas.
While the ESA may provide less
protection to species under the
jurisdiction of other countries, these
differences in ESA protections apply for
any sperm whale population that
spends time in waters of the United
States, including sperm whales within
the GOM because Mexican waters are
also outside of U.S. jurisdiction.
Therefore, we cannot rely on differences
in ESA protections for sperm whales
within the GOM and outside of the
GOM as support for the discreteness
criterion of the DPS policy.
With regard to other regulatory
mechanisms, the United States and
Mexico are both parties to the
Convention on the International Trade
in Endangered Species of Wild Fauna
and Flora (CITES), and the sperm whale
is listed on Cites Appendix I, which
means, aside from exceptional
circumstances, commercial trade of
products of sperm whales across
international borders of member
countries is prohibited. However, many
other countries within the range of the
sperm whale are parties to CITES and,
therefore, are subject to the same
prohibitions. The United States and
Mexico are also members of the
International Whaling Commission
(IWC) and have therefore adopted the
IWC’s General Principles for
Whalewatching, which include:
Managing the development of
whalewatching to minimize the risk of
adverse impacts; designing,
maintaining, and operating platforms to
minimize the risk of adverse impacts on
cetaceans, including disturbance from
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noise; and allowing the cetaceans to
control the nature and duration of
interactions. But again, many other
countries are members of the IWC, too.
We find that regulatory mechanisms
with respect to sperm whales in the
GOM do not differ significantly from
regulatory mechanisms with respect to
other sperm whale populations.
Therefore, we find that the GOM
population is not discrete from other
populations of the sperm whale based
on differences in control of exploitation,
management of habitat, conservation
status, or regulatory mechanisms.
Relation between ‘‘stock’’ and DPS—
NMFS has identified the Northern Gulf
of Mexico sperm whale population as a
stock for purposes of the Marine
Mammal Protection Act (MMPA)
https://www.nmfs.noaa.gov/pr/pdfs/sars/
ao2012whsp-gmxn.pdf (Waring et al.
(2012)). However, a stock under the
MMPA is not equivalent to a DPS under
the ESA. Under the MMPA, a
‘‘population stock’’ or ‘‘stock’’ is ‘‘a
group of marine mammals of the same
species or smaller taxa in a common
spatial arrangement that interbreed
when mature’’ (16 U.S.C. 1362(11)). The
term ‘‘stock’’ is interpreted consistent
with Congressional findings and policy:
‘‘. . . the primary objective of their
management [of stocks] should be to
maintain the health and stability of the
marine ecosystem. Whenever consistent
with this primary objective, it should be
the goal to obtain an optimum
sustainable population keeping in mind
the carrying capacity of the habitat.’’ 16.
U.S.C. 1361(5). The guidelines for
preparing stock assessment reports
under the MMPA include guidelines for
identifying stocks, and they note that
ideally, a stock would be a management
unit that identifies a demographically
isolated biological population (NMFS,
2005). Demographic isolation means
that the population dynamics of the
affected group are more a consequence
of births and deaths within the group
(internal dynamics) rather than
immigration or emigration (external
dynamics) (NMFS, 2005, https://
www.nmfs.noaa.gov/pr/pdfs/sars/
gamms2005.pdf). A major goal of
identifying stocks under the guidelines
is to avoid potential for localized
depletion where marine mammals are
subject to human-caused mortality and
serious injury.
As described above, our joint
USFWS–NMFS DPS policy contains
different criteria for identifying a
population as a DPS. The ESA’s purpose
of providing for the conservation of
species and the ecosystems upon which
they depend, along with the
Congressional direction to use the
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Federal Register / Vol. 78, No. 219 / Wednesday, November 13, 2013 / Notices
provision sparingly, guided the
development of the DPS policy. The
DPS policy requires that a population be
both discrete from other populations
and significant to the taxon to which it
belongs. While in most circumstances
we evaluate some or all of the same
evidence in determining whether a
population of marine mammals should
be considered a stock under the MMPA
or a DPS for purposes of the ESA,
demographic independence alone does
not suffice to establish a DPS. Therefore,
the fact that the GOM population is
considered a stock under the MMPA
does not qualify the population as a DPS
under the ESA.
In the 2006 NMFS Workshop on
Conservation Units of Managed Fish,
Threatened or Endangered Species, and
Marine Mammals (NOAA Tech Memo
NMFS–OPR–37, 2008), NMFS
elaborated on the distinctions:
‘‘Conservation units under the ESA
should be substantially reproductively
isolated from one another to be listed
under this act. On the other hand,
objectives of the MMPA include keeping
populations or stocks of animals above
their Optimum Sustainable Populations
OSP levels. The Magnuson-Stevens Act
(MSA) allows for management units that
may contain multiple species as
members of a complex, but the concept
of demographically independent stocks
within a species is commonly used to
determine the status of fishery
resources. Thus, demographic
independence is an appropriate basis for
identifying conservation units
(distinguishing among populations or
stocks) for the MSA and MMPA.’’
‘‘A low amount of exchange among
groups for breeding may be sufficient to
prevent development of important
genetic differences; however, these
groups may remain demographically
independent from one another.
Therefore, it is generally expected that
conservation units identified on the
basis of reproductive isolation would be
larger than those identified on the basis
of demographic independence. Thus,
discrete groups under the DPS policy
would generally be larger than discrete
groups identified for management under
the MSA or MMPA. Furthermore,
marine mammal biology includes
internal fertilization, live birth, parental
care, and maintenance of family groups;
these features act as barriers to mixing
among groups and help produce finescale population structure.’’
While Waring et al. (2012) note that
results of multi-disciplinary research
conducted in the GOM since 2000
confirm speculation by Schmidly (1981)
and indicate that GOM sperm whales
constitute a stock that is distinct from
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other Atlantic Ocean stocks(s) (Mullin et
al. 2003; Jaquet 2006; Jochens et al.
2008), it is important to note that
Waring et al. (2012) is a stock
assessment conducted under the
MMPA. A conclusion that northern
GOM sperm whales constitute a stock
under the MMPA does not demonstrate
that the GOM population of sperm
whales is a DPS.
Recovery Plan and DPSs—Our
Recovery Plan (NMFS, 2010a) and 5year review of the sperm whale (NMFS,
2009) recognize that there may be
potential sperm whale DPSs, but they
also state that further information is
needed to determine a global DPS
structure. Further, the Recovery Plan
did not use the criteria in the DPS
policy when making its assertion.
Neither document concluded that at this
time sufficient evidence exists to
identify any population as a DPS under
the ESA. Further information to support
this is not available.
DPS Analysis—Discreteness Conclusion
To summarize, the best available
information on genetics, size, behavior,
and regulatory mechanisms does not
indicate the sperm whales in the GOM
are discrete from other populations of
the sperm whale. The weight of the
evidence does not indicate the GOM
population of the sperm whale is
‘‘markedly separated’’ from other
populations. While mtDNA analysis
indicates some population structuring,
nDNA analysis indicates that successful
reproductive-mixing is occurring and
that the GOM sperm whales are not
reproductively isolated. Average size of
the individuals and number in a group
may differ throughout the range, but this
does not indicate ‘‘marked’’ differences
between sperm whales in the GOM and
sperm whales in other geographic areas.
With regard to behavioral differences,
there is evidence that sperm whales in
the GOM may use different codas for
communication, but this differentiation
is also seen within and between smaller
social groups. We found that regulatory
mechanisms with regard to sperm
whales in the GOM do not differ
significantly from those with regard to
sperm whales in other areas. We believe
the best available scientific and
commercial information does not show
that GOM sperm whales are ‘‘markedly’’
separated from other sperm whales as a
consequence of physical, physiological,
ecological, or behavioral factors.
Conclusion Regarding DPS
On the basis of the best available
information, as described above, we
conclude the GOM population is not
discrete from other sperm whale
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populations and therefore does not meet
the DPS criteria. Because the GOM
sperm whales are not discrete from
other sperm whale populations, we do
not need to determine whether the GOM
population of the sperm whale is
significant to the global taxon of sperm
whale, per the DPS policy. In any event,
even if the GOM population of the
sperm whale qualified as a discrete
population, it does not meet the
significance criterion of the DPS policy.
It does not persist in an ecological
setting unusual or unique for the taxon,
as there are other areas within the range
of the sperm whale with similar features
to the GOM (e.g., Mediterranean Sea,
which is another semi-enclosed,
partially land-locked, intercontinental,
marginal sea (www.gulfmex.org/aboutthe-gulf/gulf-of-mexico-facts/).
Loss of the GOM population would
not result in a significant gap in the
range of the sperm whale, as the range
of the GOM population (1,500,000 sq
km, www.gulfbase.org/facts.php—
visited on September 27, 2013) is only
a small portion (0.47 percent) of the
global range (317,453,000 sq km,
ngdc.noaa.gov/mgg/global/
etopo1_ocean_volumes.html). The GOM
population is not the only surviving
natural occurrence of the sperm whale,
as the species occurs in the Pacific,
Indian, and Atlantic oceans. Finally, as
discussed above, the GOM population
does not differ markedly from other
populations of the species in its genetic
characteristics.
Therefore, the GOM population of the
sperm whale does not qualify as a DPS.
Analysis of ESA Section 4(a)(1) Factors
Because the sperm whale population
in the GOM does not qualify as a DPS
under the ESA, we did not conduct an
inquiry of the factors identified in
Section 4(a)(1) of the ESA. The sperm
whale is currently listed globally as
endangered and receiving the full
protection of the ESA.
Finding
We find that the GOM population of
the sperm whale does not meet the DPS
Policy criteria for qualifying as a DPS.
Therefore, listing this population as a
separate DPS under the ESA is not
warranted.
Authority
The authority for this action is the
Endangered Species Act of 1973, as
amended (16 U.S.C. 1531 et seq.).
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Dated: November 6, 2013.
Samuel D. Rauch III,
Deputy Assistant Administrator for
Regulatory Programs, performing the
functions and duties of the Assistant
Administrator for Fisheries, National Marine
Fisheries Service.
[FR Doc. 2013–27180 Filed 11–12–13; 8:45 am]
DEPARTMENT OF COMMERCE
National Telecommunications and
Information Administration
Membership of the National
Telecommunications and Information
Administration’s Performance Review
Board
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and Information Administration,
Department of Commerce.
ACTION: Notice of membership on the
National Telecommunications and
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Performance Review Board
Membership.
AGENCY:
DEPARTMENT OF COMMERCE
National Oceanic and Atmospheric
Administration
Science Advisory Board (SAB)
Office of Oceanic and
Atmospheric Research (OAR), National
Oceanic and Atmospheric
Administration (NOAA), Department of
Commerce (DOC).
AGENCY:
Notice of time change and
meeting location.
ACTION:
The notice of an open meeting
of the NOAA Science Advisory Board
(SAB) was published in the Federal
Register on October 2, 2013
(78FR60851). Since the publication of
the meeting notice, the starting time for
the meeting on November 19, 2013 has
changed from 10:00 a.m. to 10:30 a.m.
and the meeting adjournment has
changed from 2:30 p.m. on November
20, 2013 to 11:30 a.m. The meeting will
be held at the Beacon Hotel, 1615 Rhode
Island Avenue, Washington, DC 20036.
Please see the Web site, https://
www.sab.noaa.gov/Meetings/
meetings.html for the most recent
agenda and directions to the meeting
location.
SUMMARY:
Dr.
Cynthia Decker, Executive Director,
Science Advisory Board, NOAA, Rm.
11230, 1315 East-West Highway, Silver
Spring, Maryland 20910. (Phone: 301–
734–1156, Fax: 301–713–1459. Email:
Cynthia.Decker@noaa.gov; or visit the
NOAA SAB Web site at https://
www.sab.noaa.gov.
FOR FURTHER INFORMATION CONTACT:
sroberts on DSK5SPTVN1PROD with NOTICES
SUPPLEMENTARY INFORMATION:
None.
Dated: November 6, 2013.
Jamie Krauk,
Acting Chief Financial Officer/Chief
Administrative Officer, Office of Oceanic and
Atmospheric Research, National Oceanic and
Atmospheric Administration.
[FR Doc. 2013–27178 Filed 11–12–13; 8:45 am]
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In accordance with 5 U.S.C.
4314(c)(4), the National
Telecommunications and Information
Administration (NTIA), Department of
Commerce (DOC), announce the
appointment of those individuals who
have been selected to serve as members
of NTIA’s Performance Review Board.
The Performance Review Board is
responsible for (1) reviewing
performance appraisals and rating of
Senior Executive Service (SES) members
and (2) making recommendations to the
appointing authority on other
performance management issues, such
as pay adjustments, bonuses and
Presidential Rank Awards for SES
members. The appointment of these
members to the Performance Review
Board will be for a period of twenty-four
(24) months.
DATES: The period of appointment for
those individuals selected for NTIA’s
Performance Review Board begins on
November 13, 2013.
FOR FURTHER INFORMATION CONTACT:
Ruthie B. Stewart, Department of
Commerce, Office of Human Resources
Management, Office of Executive
Resources, 14th and Constitution
Avenue NW., Room 51010, Washington,
DC 20230, at (202) 482–3130.
SUPPLEMENTARY INFORMATION: In
accordance with 5 U.S.C. 4314 (c) (4),
the National Telecommunications and
Information Administration (NTIA),
Department of Commerce (DOC),
announce the appointment of those
individuals who have been selected to
serve as members of NTIA’s
Performance Review Board. The
Performance Review Board is
responsible for (1) reviewing
performance appraisals and rating of
Senior Executive Service (SES) members
and (2) making recommendations to the
appointing authority on other
performance management issues, such
as pay adjustments, bonuses and
Presidential Rank Awards for SES
SUMMARY:
PO 00000
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68037
members. The appointment of these
members to the Performance Review
Board will be for a period of twenty-four
(24) months.
DATES: The period of appointment for
those individuals selected for NTIA’s
Performance Review Board begins on
November 13, 2013. The name, position
title, and type of appointment of each
member of NTIA’s Performance Review
Board are set forth below by
organization:
Department of Commerce, National
Telecommunications and Information
Administration (NTIA)
Fiona M. Alexander, Associate
Administrator, Office of International
Affairs, Career SES
Leonard M. Bechtel, Chief Financial
Officer and Director of
Administration, Career SES,
Chairperson
Karl B. Nebbia, Associate Administrator
for Spectrum Management, Career
SES
Alan W. Vincent, Associate
Administrator for Telecommunication
Sciences and Director Institute for
Telecommunication Sciences, Career
SES
Department of Commerce, International
Trade Administration (ITA)
Renee A. Macklin, Chief Information
Officer, Career SES
Department of Commerce, Economic
Development Administration (EDA)
Matthew S. Erskine, Deputy Assistant
Secretary for Economic Development,
Non-Career SES, Political Advisor
Dated: November 6, 2013.
Debbie Pfaff,
Director, Office of Staffing, Recruitment and
Classification, Department of Commerce
Human Resources Operations Center.
[FR Doc. 2013–27077 Filed 11–12–13; 8:45 am]
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Patent and Trademark Office
[Docket No. PTO–P–2013–0054]
Notice of Roundtable on the Renewal
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ACTION: Notice of public meeting.
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In accordance with the
Paperwork Reduction Act (PRA), the
United States Patent and Trademark
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inviting written public comment on the
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Agencies
[Federal Register Volume 78, Number 219 (Wednesday, November 13, 2013)]
[Notices]
[Pages 68032-68037]
From the Federal Register Online via the Government Printing Office [www.gpo.gov]
[FR Doc No: 2013-27180]
-----------------------------------------------------------------------
DEPARTMENT OF COMMERCE
National Oceanic and Atmospheric Administration
[Docket No. 1206013325-3912-03]
RIN 0648-XA983
Endangered and Threatened Wildlife; Notice of 12-Month Finding on
a Petition To List the Sperm Whale (Physeter macrocephalus) as an
Endangered or Threatened Distinct Population Segment (DPS) in the Gulf
of Mexico
AGENCY: National Marine Fisheries Service (NMFS), National Oceanic and
Atmospheric Administration (NOAA), Commerce.
ACTION: Status review; notice of finding.
-----------------------------------------------------------------------
SUMMARY: We, NMFS, announce a 12-month finding on a petition to list
the sperm whale (Physeter macrocephalus) in the Gulf of Mexico as an
endangered or threatened distinct population segment (DPS) under the
Endangered Species Act of 1973 as amended (ESA). We conducted a review
of the status of this population, as described below. Based on the best
available scientific and commercial information, we find that the
petitioned action is not warranted.
DATES: The finding announced in this notice was made on November 13,
2013.
ADDRESSES: Information used to make this finding is available for
public inspection by appointment during normal business hours at NMFS
Headquarters, Protected Resources Office, 1315 East-West Highway,
Silver Spring, MD 20910. This file includes the information provided by
the public and scientific and commercial information gathered for the
status review. The petition and a list of the references we used can
also be found at https://www.nmfs.noaa.gov/pr/.htm.
FOR FURTHER INFORMATION CONTACT: Marta Nammack, NMFS, Office of
Protected Resources, (301) 427-8469.
SUPPLEMENTARY INFORMATION: On December 9, 2011, we received a petition
from WildEarth Guardians to list the sperm whale (Physeter
macrocephalus) population in the Gulf of Mexico as an endangered or
threatened Distinct Population Segment (DPS) under the Endangered
Species Act (ESA); sperm whales are currently listed as a single
endangered species throughout their global range (35 FR 8495; June 2,
1970). The petitioner also requested designation of critical habitat
concurrent with the listing.
After reviewing the petition, the literature cited in the petition,
and other literature and information available in our files, we found
that the petition met the requirements of the regulations under 50 CFR
424.14(b)(2) and determined that the petition presented substantial
information indicating that the petitioned action may be warranted (78
FR 19176; March 29, 2013). At that time, we commenced a status review
of the sperm whale in the Gulf of Mexico and solicited information
pertaining to the population. Section 4(b)(3)(B) of the ESA requires
that when a petition to revise the List of Endangered and Threatened
Wildlife and Plants is found to present substantial scientific and
commercial information, we make a finding on whether the petitioned
action is (a) not warranted, (b) warranted, or (c) warranted but
precluded from listing by other pending proposals of higher priority.
This finding is to be made within 12 months of the date the petition
was received, and the finding is to be published promptly in the
Federal Register.
There are two key tasks associated with conducting an ESA status
review. The first is to determine whether the petitioned entity
qualifies as one or more species under the ESA. The ESA defines the
term ``species'' to include ``any distinct population segment of any
species of vertebrate fish or wildlife which interbreeds when mature.''
If the petitioned entity qualifies as a species, the second task is to
conduct an extinction risk assessment to determine whether the species
is threatened or endangered. The ESA defines the term ``endangered
species'' as ``any species which is in danger of extinction throughout
all or a significant portion of its range.'' The term ``threatened
species'' is defined as ``any species which is likely to become
endangered within the foreseeable future throughout all or a
significant portion of its range.'' Thus, we interpret an ``endangered
species'' to be one that is presently in danger of extinction. A
``threatened species,'' on the other hand, is not presently in danger
of extinction, but is likely to become so in the foreseeable future
(that is, at a later time). In other words, the primary statutory
difference between a threatened and endangered species is the timing of
when a species may be in danger of extinction, either presently
(endangered) or in the foreseeable future (threatened).
Species Background
The sperm whale (Linnaeus, 1758) is listed as an endangered species
under the ESA. It was first listed under the precursor to the ESA, the
Endangered Species Conservation Act of 1969, and remained on the list
of threatened and
[[Page 68033]]
endangered species after the passage of the ESA in 1973 (35 FR 18319;
December 2, 1970). Whaling was the main reason for listing the sperm
whale. Commercial whaling for this species ended in 1988 with the
implementation of a moratorium against whaling by the International
Whaling Commission (IWC). While whaling was eliminated by the IWC
whaling moratorium, several potential threats remain, as discussed in
the sperm whale recovery plan (NMFS, 2010a). Sperm whales are deep and
prolonged divers and use the entire water column, even in very deep
areas. Most sperm whales are found in very deep waters (>3,000 m), but
they generally feed between 500-1,000 m where most of their prey is
found. Sperm whales feed primarily on large- and medium-sized squid,
but the list of documented food items is fairly long and diverse,
including other cephalopods and medium-and large-sized demersal fish,
such as rays, sharks, and many teleosts (Berzin, 1972; Clarke 1977,
1980; Rice, 1989). The diet of large males in some areas, especially in
high northern latitudes, is dominated by fish (Rice, 1989). Lockyer
(1981) estimated sperm whales consumed about 3.0-3.5 percent of their
body weight per day.
Sperm whales are perhaps the most widely distributed mammal species
on Earth. The social organization of most mammals is characterized by
female philopatry and male dispersal. Groups of females and juveniles
are found mainly at low latitudes, while males reach polar waters,
returning to tropical and subtropical waters to breed. Sperm whales are
organized in groups in which females (some related to each other and
some not) travel with their sub-adult offspring. Mature female and
immature sperm whales of both sexes are found in more temperate and
tropical waters from the equator to around 45[ordm]N throughout the
year. Adult males will move extensively, even to polar waters, and then
return to tropical and subtropical waters.
Sperm whales mature slowly and can live to ages in excess of 60
years (Rice, 1989). Females usually begin ovulating at 7-13 years of
age and usually conceive at about age 9 (Rice, 1989). Maturation in
males usually begins in this same age interval, but most individuals do
not become fully mature until their twenties. In the North Atlantic
Ocean, the peak breeding season for sperm whales occurs during the
spring (March/April to June), although some mating activity occurs
December to August. In the South Atlantic the peak breeding season is
presumed to occur in the austral spring. During mating seasons, prime
bulls in their late twenties and older rove among groups of females.
Because females within a group often come into estrus synchronously,
the males need not remain with the females for the breeding season to
achieve maximal breeding success (Best and Butterworth, 1980) and their
association with a group can be as brief as several hours. Gestation
lasts well over a year, with credible estimates of the normal duration
ranging from 15 months to more than a year and a half. Lactation lasts
at least 2 years, and the inter-birth-interval is 4-6 years (Best et
al., 1984) for prime-aged females. Female sperm whales rarely become
pregnant after the age of 40 (Whitehead, 2003). Two particular aspects
of the sperm whale's reproductive biology are relevant to recovery.
First, the maximal rate of increase in reproduction is very low,
perhaps no more than one or two percent per year. Second, selective
killing of large males by modern whaling could have had the residual
effect of reducing reproductive rates (Whitehead et al., 1997).
Status Review
Our 90-day finding accepting the petition solicited information
from the public and initiated a status review of the sperm whale in the
Gulf of Mexico (GOM) to gather any additional information to inform our
review of the petitioned action and our application of the DPS policy.
We reviewed the best available information, and we conducted a DPS
analysis to determine whether the GOM population of the sperm whale
qualifies as a DPS under the ESA. Here we review the best available
information on physical, physiological, ecological, and behavioral
factors to determine whether the GOM population is discrete.
Are sperm whales in the Gulf of Mexico discrete from other sperm whale
populations?
The ESA provides for listing species, subspecies, or DPSs of
vertebrate species. When we evaluate a petition to list an entity as
threatened or endangered under the ESA, we must first determine whether
the petitioned entity qualifies as a species under the ESA. This
petition argues that the Gulf of Mexico sperm whale population meets
the requirements for being identified as a DPS and requests we list
sperm whales in the Gulf of Mexico as a threatened or endangered DPS.
Our joint NMFS-U.S. Fish and Wildlife Service (USFWS) Policy on
Recognition of Distinct Vertebrate Population Segments under the
Endangered Species Act (DPS policy) (61 FR 4722; February 7, 1996)
identifies two elements that must be considered when identifying a DPS:
(1) The discreteness of the population segment in relation to the
remainder of the species (or subspecies) to which it belongs; and (2)
the significance of the population segment to the species to which it
belongs. A population segment of a vertebrate species may be considered
discrete if it satisfies either one of the following conditions: (1) It
is markedly separated from other populations of the same taxon as a
consequence of physical, physiological, ecological, or behavioral
factors. Quantitative measures of genetic or morphological
discontinuity may provide evidence of this separation; or (2) it is
delimited by international governmental boundaries within which
differences in control of exploitation, management of habitat,
conservation status, or regulatory mechanisms exist that are
significant in light of section 4(a)(1)(D) of the ESA. If a population
segment is considered discrete by one or more of the above conditions,
its biological and ecological significance will then be considered in
light of Congressional guidance (see Senate Report 151, 96th Congress,
1st Session) that the authority to list DPSs be used ``. . .
sparingly'' while encouraging the conservation of genetic diversity.
The DPS policy directs us to consider available scientific evidence of
the discrete population segment's importance to the taxon to which it
belongs. This consideration may include, but is not limited to, the
following: (1) Persistence of the discrete population segment in an
ecological setting unusual or unique for the taxon; (2) evidence that
loss of the discrete population segment would result in a significant
gap in the range of a taxon; (3) evidence that the discrete population
segment represents the only surviving natural occurrence of a taxon
that may be more abundant elsewhere as an introduced population outside
its historic range; or (4) evidence that the discrete population
segment differs markedly from other populations of the species in its
genetic characteristics.
DPS Analysis
To determine if the sperm whale in the GOM meets the DPS criteria,
we evaluate the best available information to determine whether sperm
whales in the Gulf of Mexico are markedly separated as a consequence of
physical, physiological, ecological, or behavioral factors from other
populations of the sperm whale.
[[Page 68034]]
Genetics--An examination of the best available genetic information
reveals that, although there is strong mtDNA evidence of population
structuring indicating differences between the GOM population and sperm
whales in the northwest Atlantic, this is not coupled with nDNA
evidence that would indicate that males from the GOM are genetically
different from males in the northwest Atlantic. Physically mature male
sperm whales typically range over huge distances on their own (Best,
1979; Rice, 1989; Whitehead, 1993; Whitehead and Weilgart, 2000; Teloni
et al., 2008). In contrast to females, males disperse from their natal
units at a mean estimated age of 6 years, when they migrate slowly into
higher latitudes prior to attaining sexual maturity at 18-21 years
(Whitehead and Weilgart, 2000). This is re[fllig]ected in high
variability and a lack of geographical structure in nDNA relative to
mtDNA (Lyrholm et al., 1999).
There are statistically significant patterns of mtDNA
differentiation between oceans (Engelhaupt, 2004; SWSS, 2008;
Engelhaupt et al., 2009; NMFS, 2010a); however, studies examining nDNA
reveal either no significant (Lyrholm et al., 1999) or low (Bond, 1999)
degrees of population structuring between oceans. Engelhaupt et al.
(2009) suggest that the discrepancy between mtDNA and nDNA
differentiation may reflect sex biased dispersal, and male mediated
gene flow may connect geographically isolated regions on an oceanic
scale. Their analysis of nDNA showed no significant difference between
whales sampled in the GOM and those from other areas of the North
Atlantic, indicating that mature males move in and out of the GOM. The
results of the Engelhaupt et al. (2009) study indicate that population
structuring is different for mtDNA compared with population structuring
for nDNA.
At best, mtDNA evidence suggests that females are philopatric;
however, mtDNA does not alone describe population structure. Because
mtDNA is maternally inherited, differences in mtDNA haplotypes between
populations do not necessarily mean that the populations are
substantially reproductively isolated from each other because they do
not provide any information on males. Due to the wide ranging nature of
mature male sperm whales, males from one population may breed with
females from other populations. We have indicated in other status
reviews that mtDNA data may indicate that populations are discrete, but
in species where female and male movement patterns differ, nDNA data
may indicate that the populations are homogeneous (see e.g., loggerhead
sea turtle, 68 FR 53947, September 15, 2003 at 53950-51 and Conant et
al., 2009, at 18, 22, 25-28; southern resident killer whale, Krahn et
al., 2002, at 23-30). As noted in SWSS (2008), a male sperm whale
tagged in 2002 moved into the North Atlantic for more than 2 months,
providing the first evidence that the GOM population may not be a stock
isolated from the North Atlantic (SWSS, 2008; Waring et al., 2012). Its
return to the GOM included an extended stay off the northwest Cuban
coast, and it summered in two different regions of the upper GOM and
visited the Gulf of Campeche twice (SWSS, 2008). While some may view
this as support for separate stocks in the GOM and the North Atlantic,
SWSS (2008) notes that few males were sampled in the GOM. Because the
tags were deployed from June to early August, more individuals were
tracked during the summer months (SWSS, 2008). Therefore, it is likely
that mature males were not in the GOM at this time, as they spend most
of their time in colder waters at high latitudes and only visit
tropical waters to reproduce (Best 1979; Whitehead and Arnbom 1987;
Whitehead 2003, as cited in SWSS (2008)).
The fact that males move in and out of the GOM and interbreed with
females from other populations when mature, as evidenced by the
homogeneity of the nDNA, indicates that the GOM population is not
markedly separated from other populations in the Atlantic Ocean.
Engelhaupt et al. (2009) demonstrate that a single, undivided genetic
population of sperm whales is found from the GOM to at least northern
Europe. As we have summarized here, the best available genetic
information indicates that sperm whales in the GOM are not discrete
from other sperm whale populations.
Vocalization--We next examined information on codas. Sperm whale
social structure is complex, with females, calves, and immature animals
of both sexes living in relatively stable social ``units'' containing
on average 11-12 animals that persist for decades (Rendell and
Whitehead, 2004). These sperm whale social groups communicate via
codas: Repeated stereotyped sequences of 3-40 broadband (0-16 kHz)
clicks generally heard during periods of socializing (Watkins and
Schevill, 1977). Codas are shared among individuals of a social unit
and are considered to be primarily for intra-group communication
(Weilgart and Whitehead, 1997; Rendell and Whitehead, 2004). These
distinctive, short, patterned series of clicks are associated with
social behavior and interactions within social groups (Weilgart and
Whitehead, 1993).
Significant differences in vocalization or coda repertoire exist
amongst smaller social groups or ``units'' of sperm whales, and this
variation amongst social units or groups is commonplace for sperm
whales (Weilgart and Whitehead, 1997; Rendell and Whitehead, 2004).
Differences in vocalization are culturally transmitted by the
matrilineal line, and there is a difference between geographical sperm
whale variation in codas (macrogeographic) and coda ``dialects''
(microgeographic) (Mundinger, 1982). In a study of sperm whales in the
southern Pacific Ocean, Weilgart and Whitehead (1997) found that the
sperm whale groups they encountered had distinctive dialects in coda
usage based on analyses of interclick intervals (ICIs), the time
intervals between clicks in a coda, standardized to total coda length.
The group-specific dialects that are found in sperm whales have even
been deemed as similar to those which occur in killer whale ``vocal
clans'' (Weilgart and Whitehead, 1997; Rendell and Whitehead, 2003).
Codas and mtDNA have been linked; a study of six sperm whale groups
revealed a clear link between mtDNA and coda repertoire as groups with
similar mtDNA tended to have similar coda usage dialects (Whitehead et
al., 1998). These results indicate codas are transmitted across
generations matrilineally. Whitehead et al. (1998) suggested vertical
cultural transmission (offspring learn codas from their mothers) as the
best explanation for this pattern. This may reflect the mtDNA
information presented above suggesting population structure, without
consideration of the nDNA. The sperm whale seismic study (SWSS, 2008)
cited in the petition found variation in vocalization between the north
central GOM and the northwest GOM. Because there is evidence of
different types of coda variation (i.e., macrogeographic versus
microgeographic dialects) within the GOM, communication is passed down
from the mother, and adult male sperm whales travel outside the Gulf of
Mexico, the communication difference between GOM sperm whales and sperm
whales from other populations does not indicate sperm whales in the GOM
are ``markedly'' separate.
Group size--While group size in the GOM is smaller on average than
in other oceans, group size is variable throughout their global range.
The fact that group sizes are similar to those in the Caribbean and
smaller than group sizes in some other oceans (SWSS,
[[Page 68035]]
2008) does not show a ``marked'' separation from other sperm whale
group sizes. Christal et al. (1998) note that estimated social unit
size in the Galapagos, for example, ranged from 3 to 24 individuals and
presented evidence of splitting and merging of units and of transfer of
individuals between units. The considerable variation in unit size
(perhaps caused by demographic processes) suggests that the benefits of
remaining in a social unit usually outweigh selection for some optimal
unit size (Christal et al., 1998). Richter et al. (2008) note that it
could be argued that differences in ecological conditions in which
various sperm whale populations live are reflected in the parameters of
their social behavior, such as group size and association rate (Richter
et al., 2008). The best available evidence does not indicate that sperm
whale group size in the GOM is different from all other populations of
the sperm whale.
Whale size--Mean size of sperm whales in the GOM (8.5 m) has been
reported to be smaller than that of other sperm whale populations
(e.g., 10 m for the Gulf of California population) (SWSS, 2008). While
photographic data on known males and sound pulse studies showed that
those measured in the GOM were smaller than breeding males elsewhere
(Jaquet et al., 2006; Antunes et al., 2006), no mature males have been
observed in the GOM. This only confirms that younger male whales that
have recently departed from their mothers are smaller than those at
full maturity, which is not noteworthy. Older males, which apparently
only pass through the GOM for breeding, are larger than the younger
males that have not yet migrated out of the GOM. Further, whale size
data from these studies have never been normalized to account for age,
so a reliable comparison cannot be made. Finally, Jochens et al. (2008)
argue that female/adolescent size differences among sperm whale
populations may be the result of nothing more than differences in prey,
suggesting that ``it is possible that the population studied is smaller
because smaller animals may prefer the shallower waters relative to
their diving ability and/or availability of suitable prey.'' Whales may
assort themselves by water depths to match their body sizes. Finally,
even if GOM whales are a little smaller on average than other
populations of sperm whale, such a modest difference is not sufficient
to demonstrate that the GOM population is ``markedly separated'' from
other sperm whale populations.
International boundaries--In examining whether a population is
discrete based on international governmental boundaries, we are to
examine differences in the control of exploitation, management of
habitat, conservation status, or regulatory mechanisms exist that are
significant in light of section 4(a)(1)(D) of the ESA. Section
4(a)(1)(D), the inadequacy of existing regulatory mechanisms, is one of
the five factors we must evaluate to determine whether to list a
species. We did not find any information pointing to significant
differences in control of exploitation, management of habitat,
conservation status, or regulatory mechanisms between the population of
sperm whales in the GOM and any other particular population of the
sperm whale such that the population of the sperm whale in the GOM
could be considered discrete from a sperm whale population outside of
the GOM. The ESA extends prohibitions against take of endangered
species by any person subject to the jurisdiction of the United States
within the United States, its territorial waters, or on the high seas.
While the ESA may provide less protection to species under the
jurisdiction of other countries, these differences in ESA protections
apply for any sperm whale population that spends time in waters of the
United States, including sperm whales within the GOM because Mexican
waters are also outside of U.S. jurisdiction. Therefore, we cannot rely
on differences in ESA protections for sperm whales within the GOM and
outside of the GOM as support for the discreteness criterion of the DPS
policy.
With regard to other regulatory mechanisms, the United States and
Mexico are both parties to the Convention on the International Trade in
Endangered Species of Wild Fauna and Flora (CITES), and the sperm whale
is listed on Cites Appendix I, which means, aside from exceptional
circumstances, commercial trade of products of sperm whales across
international borders of member countries is prohibited. However, many
other countries within the range of the sperm whale are parties to
CITES and, therefore, are subject to the same prohibitions. The United
States and Mexico are also members of the International Whaling
Commission (IWC) and have therefore adopted the IWC's General
Principles for Whalewatching, which include: Managing the development
of whalewatching to minimize the risk of adverse impacts; designing,
maintaining, and operating platforms to minimize the risk of adverse
impacts on cetaceans, including disturbance from noise; and allowing
the cetaceans to control the nature and duration of interactions. But
again, many other countries are members of the IWC, too. We find that
regulatory mechanisms with respect to sperm whales in the GOM do not
differ significantly from regulatory mechanisms with respect to other
sperm whale populations. Therefore, we find that the GOM population is
not discrete from other populations of the sperm whale based on
differences in control of exploitation, management of habitat,
conservation status, or regulatory mechanisms.
Relation between ``stock'' and DPS--NMFS has identified the
Northern Gulf of Mexico sperm whale population as a stock for purposes
of the Marine Mammal Protection Act (MMPA) https://www.nmfs.noaa.gov/pr/pdfs/sars/ao2012whsp-gmxn.pdf (Waring et al. (2012)). However, a stock
under the MMPA is not equivalent to a DPS under the ESA. Under the
MMPA, a ``population stock'' or ``stock'' is ``a group of marine
mammals of the same species or smaller taxa in a common spatial
arrangement that interbreed when mature'' (16 U.S.C. 1362(11)). The
term ``stock'' is interpreted consistent with Congressional findings
and policy: ``. . . the primary objective of their management [of
stocks] should be to maintain the health and stability of the marine
ecosystem. Whenever consistent with this primary objective, it should
be the goal to obtain an optimum sustainable population keeping in mind
the carrying capacity of the habitat.'' 16. U.S.C. 1361(5). The
guidelines for preparing stock assessment reports under the MMPA
include guidelines for identifying stocks, and they note that ideally,
a stock would be a management unit that identifies a demographically
isolated biological population (NMFS, 2005). Demographic isolation
means that the population dynamics of the affected group are more a
consequence of births and deaths within the group (internal dynamics)
rather than immigration or emigration (external dynamics) (NMFS, 2005,
https://www.nmfs.noaa.gov/pr/pdfs/sars/gamms2005.pdf). A major goal of
identifying stocks under the guidelines is to avoid potential for
localized depletion where marine mammals are subject to human-caused
mortality and serious injury.
As described above, our joint USFWS-NMFS DPS policy contains
different criteria for identifying a population as a DPS. The ESA's
purpose of providing for the conservation of species and the ecosystems
upon which they depend, along with the Congressional direction to use
the
[[Page 68036]]
provision sparingly, guided the development of the DPS policy. The DPS
policy requires that a population be both discrete from other
populations and significant to the taxon to which it belongs. While in
most circumstances we evaluate some or all of the same evidence in
determining whether a population of marine mammals should be considered
a stock under the MMPA or a DPS for purposes of the ESA, demographic
independence alone does not suffice to establish a DPS. Therefore, the
fact that the GOM population is considered a stock under the MMPA does
not qualify the population as a DPS under the ESA.
In the 2006 NMFS Workshop on Conservation Units of Managed Fish,
Threatened or Endangered Species, and Marine Mammals (NOAA Tech Memo
NMFS-OPR-37, 2008), NMFS elaborated on the distinctions:
``Conservation units under the ESA should be substantially
reproductively isolated from one another to be listed under this act.
On the other hand, objectives of the MMPA include keeping populations
or stocks of animals above their Optimum Sustainable Populations OSP
levels. The Magnuson-Stevens Act (MSA) allows for management units that
may contain multiple species as members of a complex, but the concept
of demographically independent stocks within a species is commonly used
to determine the status of fishery resources. Thus, demographic
independence is an appropriate basis for identifying conservation units
(distinguishing among populations or stocks) for the MSA and MMPA.''
``A low amount of exchange among groups for breeding may be
sufficient to prevent development of important genetic differences;
however, these groups may remain demographically independent from one
another. Therefore, it is generally expected that conservation units
identified on the basis of reproductive isolation would be larger than
those identified on the basis of demographic independence. Thus,
discrete groups under the DPS policy would generally be larger than
discrete groups identified for management under the MSA or MMPA.
Furthermore, marine mammal biology includes internal fertilization,
live birth, parental care, and maintenance of family groups; these
features act as barriers to mixing among groups and help produce fine-
scale population structure.''
While Waring et al. (2012) note that results of multi-disciplinary
research conducted in the GOM since 2000 confirm speculation by
Schmidly (1981) and indicate that GOM sperm whales constitute a stock
that is distinct from other Atlantic Ocean stocks(s) (Mullin et al.
2003; Jaquet 2006; Jochens et al. 2008), it is important to note that
Waring et al. (2012) is a stock assessment conducted under the MMPA. A
conclusion that northern GOM sperm whales constitute a stock under the
MMPA does not demonstrate that the GOM population of sperm whales is a
DPS.
Recovery Plan and DPSs--Our Recovery Plan (NMFS, 2010a) and 5-year
review of the sperm whale (NMFS, 2009) recognize that there may be
potential sperm whale DPSs, but they also state that further
information is needed to determine a global DPS structure. Further, the
Recovery Plan did not use the criteria in the DPS policy when making
its assertion. Neither document concluded that at this time sufficient
evidence exists to identify any population as a DPS under the ESA.
Further information to support this is not available.
DPS Analysis--Discreteness Conclusion
To summarize, the best available information on genetics, size,
behavior, and regulatory mechanisms does not indicate the sperm whales
in the GOM are discrete from other populations of the sperm whale. The
weight of the evidence does not indicate the GOM population of the
sperm whale is ``markedly separated'' from other populations. While
mtDNA analysis indicates some population structuring, nDNA analysis
indicates that successful reproductive-mixing is occurring and that the
GOM sperm whales are not reproductively isolated. Average size of the
individuals and number in a group may differ throughout the range, but
this does not indicate ``marked'' differences between sperm whales in
the GOM and sperm whales in other geographic areas. With regard to
behavioral differences, there is evidence that sperm whales in the GOM
may use different codas for communication, but this differentiation is
also seen within and between smaller social groups. We found that
regulatory mechanisms with regard to sperm whales in the GOM do not
differ significantly from those with regard to sperm whales in other
areas. We believe the best available scientific and commercial
information does not show that GOM sperm whales are ``markedly''
separated from other sperm whales as a consequence of physical,
physiological, ecological, or behavioral factors.
Conclusion Regarding DPS
On the basis of the best available information, as described above,
we conclude the GOM population is not discrete from other sperm whale
populations and therefore does not meet the DPS criteria. Because the
GOM sperm whales are not discrete from other sperm whale populations,
we do not need to determine whether the GOM population of the sperm
whale is significant to the global taxon of sperm whale, per the DPS
policy. In any event, even if the GOM population of the sperm whale
qualified as a discrete population, it does not meet the significance
criterion of the DPS policy. It does not persist in an ecological
setting unusual or unique for the taxon, as there are other areas
within the range of the sperm whale with similar features to the GOM
(e.g., Mediterranean Sea, which is another semi-enclosed, partially
land-locked, intercontinental, marginal sea (www.gulfmex.org/about-the-gulf/gulf-of-mexico-facts/).
Loss of the GOM population would not result in a significant gap in
the range of the sperm whale, as the range of the GOM population
(1,500,000 sq km, www.gulfbase.org/facts.php--visited on September 27,
2013) is only a small portion (0.47 percent) of the global range
(317,453,000 sq km, ngdc.noaa.gov/mgg/global/etopo1_ocean_volumes.html). The GOM population is not the only surviving natural
occurrence of the sperm whale, as the species occurs in the Pacific,
Indian, and Atlantic oceans. Finally, as discussed above, the GOM
population does not differ markedly from other populations of the
species in its genetic characteristics.
Therefore, the GOM population of the sperm whale does not qualify
as a DPS.
Analysis of ESA Section 4(a)(1) Factors
Because the sperm whale population in the GOM does not qualify as a
DPS under the ESA, we did not conduct an inquiry of the factors
identified in Section 4(a)(1) of the ESA. The sperm whale is currently
listed globally as endangered and receiving the full protection of the
ESA.
Finding
We find that the GOM population of the sperm whale does not meet
the DPS Policy criteria for qualifying as a DPS. Therefore, listing
this population as a separate DPS under the ESA is not warranted.
Authority
The authority for this action is the Endangered Species Act of
1973, as amended (16 U.S.C. 1531 et seq.).
[[Page 68037]]
Dated: November 6, 2013.
Samuel D. Rauch III,
Deputy Assistant Administrator for Regulatory Programs, performing the
functions and duties of the Assistant Administrator for Fisheries,
National Marine Fisheries Service.
[FR Doc. 2013-27180 Filed 11-12-13; 8:45 am]
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