Endangered and Threatened Wildlife and Plants; Petition To List Grand Canyon Cave Pseudoscorpion, 42654-42658 [2011-17864]
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Federal Register / Vol. 76, No. 138 / Tuesday, July 19, 2011 / Proposed Rules
ACTIONS FUNDED IN FY 2010 AND FY 2011 BUT NOT YET COMPLETED—Continued
Species
Action
2 TN River mussels (fluted kidneyshell (LPN = 2), slabside pearlymussel (LPN = 2) 5 ..........................................
Jemez Mountain salamander (LPN = 2) 5 .................................................................................................................
Proposed listing.
Proposed listing.
1 Funds
for listing actions for these species were provided in previous FYs.
funds for these high-priority listing actions were provided in FY 2008 or 2009, due to the complexity of these actions and competing
priorities, these actions are still being developed.
3 Partially funded with FY 2010 funds and FY 2011 funds.
4 Funded with FY 2010 funds.
5 Funded with FY 2011 funds.
2 Although
We have endeavored to make our
listing actions as efficient and timely as
possible, given the requirements of the
relevant law and regulations, and
constraints relating to workload and
personnel. We are continually
considering ways to streamline
processes or achieve economies of scale,
such as by batching related actions
together. Given our limited budget for
implementing section 4 of the Act, these
actions described above collectively
constitute expeditious progress.
Pinus albicaulis will be added to the
list of candidate species upon
publication of this 12-month finding.
We will continue to evaluate this
species as new information becomes
available. Continuing review will
determine if a change in status is
warranted, including the need to make
prompt use of emergency listing
procedures.
We intend that any proposed listing
determination for Pinus albicaulis will
be as accurate as possible. Therefore, we
will continue to accept additional
information and comments from all
concerned governmental agencies, the
scientific community, industry, or any
other interested party concerning this
finding.
References Cited
A complete list of references cited is
available on the Internet at https://
www.regulations.gov and upon request
from the Wyoming Ecological Services
Field Office (see ADDRESSES section).
Author(s)
The primary authors of this notice are
the staff members of the Wyoming
Ecological Services Field Office.
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Authority
The authority for this section is
section 4 of the Endangered Species Act
of 1973, as amended (16 U.S.C. 1531 et
seq.).
Dated: July 1, 2011.
Daniel M. Ashe,
Director, Fish and Wildlife Service.
[FR Doc. 2011–17943 Filed 7–18–11; 8:45 am]
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DEPARTMENT OF THE INTERIOR
Fish and Wildlife Service
50 CFR Part 17
[Docket No. FWS–R2–ES–2011–0044; MO
92210–0–0008–B2]
Endangered and Threatened Wildlife
and Plants; Petition To List Grand
Canyon Cave Pseudoscorpion
AGENCY: Fish and Wildlife Service,
Interior.
ACTION: Notice of 12-month petition
finding.
SUMMARY: We, the U.S. Fish and
Wildlife Service (Service), announce a
12-month finding on a petition to list
the Grand Canyon cave pseudoscorpion
(Archeolarca cavicola) as threatened or
endangered with critical habitat under
the Endangered Species Act of 1973, as
amended (Act). After review of the best
scientific and commercial information
available, we find that listing the Grand
Canyon cave pseudoscorpion is not
warranted at this time. However, we ask
the public to submit to us any new
information that becomes available
concerning the threats to the Grand
Canyon cave pseudoscorpion or its
habitat at any time.
DATES: The finding announced in this
document was made on July 19, 2011.
ADDRESSES: This finding is available on
the Internet at https://www.regulations.
gov at Docket Number FWS–R2–ES–
2011–0044. Supporting documentation
we used in preparing this finding is
available for public inspection, by
appointment, during normal business
hours by contacting the U.S. Fish and
Wildlife Service, Arizona Ecological
Services Field Office, 2321 W. Royal
Palm Road, Suite 103, Phoenix, AZ
85021; telephone (602) 242–0210;
facsimile (602) 242–2513. If you use a
telecommunications device for the deaf
(TDD), please call the Federal
Information Relay Service (FIRS) at
(800) 877–8339. Please submit any new
information, comments, or questions
concerning this finding to the above
address.
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FOR FURTHER INFORMATION CONTACT:
Steve Spangle, Field Supervisor, U.S.
Fish and Wildlife Service, Arizona
Ecological Services Field Office, 2321
W. Royal Palm Road, Suite 103,
Phoenix, AZ 85021; telephone (602)
242–0210; facsimile (602) 242–2513. If
you use a telecommunications device
for the deaf (TDD), please call the
Federal Information Relay Service
(FIRS) at (800) 877–8339.
SUPPLEMENTARY INFORMATION:
Background
Section 4(b)(3)(B) of the Endangered
Species Act of 1973, as amended (Act)
(16 U.S.C. 1531 et seq.), requires that,
for any petition containing substantial
scientific or commercial information
indicating that listing the species may
be warranted, we make a finding within
12 months of the date of receipt of the
petition. In this finding we determine
that the petitioned action is: (a) Not
warranted, (b) warranted, or (c)
warranted, but immediate proposal of a
regulation implementing the petitioned
action is precluded by other pending
proposals to determine whether species
are threatened or endangered, and
expeditious progress is being made to
add or remove qualified species from
the Lists of Endangered and Threatened
Wildlife and Plants. Section 4(b)(3)(C) of
the Act requires that we treat a petition
for which the requested action is found
to be warranted but precluded as though
resubmitted on the date of such finding,
that is, requiring a subsequent finding to
be made within 12 months. We must
publish these 12-month findings in the
Federal Register.
Previous Federal Actions
The Grand Canyon cave
pseudoscorpion was formerly a
candidate 2 species, a taxon for which
information in our possession indicated
that proposing to list was possibly
appropriate, but for which persuasive
data on biological vulnerability and
threats were not available to support a
proposed listing rule (54 FR 554;
January 6, 1989). The designation of
candidate 2 species was discontinued in
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1996; therefore, the species has no
candidate status.
On June 25, 2007, we received a
formal petition dated June 18, 2007,
from Forest Guardians (now WildEarth
Guardians), requesting that we do the
following: (1) Consider all full species
in our Southwest Region ranked as G1
or G1G2 by the organization
NatureServe for listing, except those
that are currently listed, proposed for
listing, or candidates; and (2) list each
species under the Act as either
endangered or threatened with critical
habitat. The petitioners presented two
tables that collectively listed 475
species for consideration and requested
that the Service incorporate all analyses,
references, and documentation provided
by NatureServe in its online database
https://www.natureserve.org/ into the
petition. The petition clearly identified
itself as a petition and included the
appropriate identification information,
as required in 50 CFR 424.14(a). We
acknowledged the receipt of the petition
in a letter to WildEarth Guardians dated
July 11, 2007.
On December 16, 2009, we made a 90day finding that the petition presented
substantial scientific information
indicating that listing 67 of the 475
species may be warranted; the Grand
Canyon cave pseudoscorpion
(incorrectly referenced as the Grand
Canyon cave scorpion) was in that
group of 67 species. Based on the
evaluation of the information provided
in the petition, we determined that the
petition presented substantial
information to indicate that listing the
Grand Canyon cave pseudoscorpion
may be warranted due to the present or
threatened destruction, modification, or
curtailment of its habitat or range
resulting from groundwater pollution
and recreational impacts, and to the
inadequacy of existing regulatory
mechanisms resulting from unregulated
visitation. The 90-day finding and our
initiation of a status review was
published in the Federal Register on
December 16, 2009 (74 FR 66866). This
notice constitutes the 12-month finding
on the June 18, 2007, petition to list the
Grand Canyon cave pseudoscorpion as
threatened or endangered.
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Species Information
Species Description
In 1978 W. Calvin Welbourn collected
one female specimen of the Grand
Canyon cave pseudoscorpion (Class
Arachnida, Order Pseudoscorpionida,
Family Garypidae) in Grand Canyon
National Park’s Cave of the Domes. This
specimen was first described by
Muchmore (1981, p. 55). Welbourn’s
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unpublished report (1978, p. 40) stated
that the specimen was an undescribed
troglophile. A troglophile is a species
that can spend its entire life within
caves, does not exhibit adaptations for
living in caves, but can also be found in
suitable habitats outside of caves, such
as mines or animal burrows. This is in
comparison to troglobites, which are
species that are found exclusively in
caves and have developed adaptations
for cave life, such as heightened sense
of hearing, touch, and smell. The Grand
Canyon cave pseudoscorpion was
recognized within the genus
Archeolarca and was given the species
name cavicola in recognition of its
subterranean habitat where it was
collected by Welbourn (1981, p. 55).
The specimen collected in 1978 is the
only one known to exist. No other
individuals are known to have been
collected since 1978, although very little
effort has been made to collect this or
other species in the genus (Service 1991,
p. 3).
Pseudoscorpions are tiny arachnids
bearing large chelae, or claws, but
lacking a telson, or stinger, that true
scorpions possess. The specimen of
Grand Canyon cave pseudoscorpion,
when compared to other species of
pseudoscorpions, such as Archeolarca
welbourni and Archeolarca
guadalupensis, was considered large
(0.12 inches (in) or 3.03 millimeters
(mm)), had longer appendages, more
reduced posterior eyes, and fewer setae
(stiff bristles present on the body) on its
upper dorsal section (Muchmore 1981,
p. 56). Muchmore (1981, pp. 52–56)
described three new species of
Archeolarca, and concluded that the
Grand Canyon cave pseudoscorpion
showed the greatest overall adaption to
the cave environment. Welbourn (1978,
p. 40) noted that the specimen appeared
to be similar to the pseudoscorpions
from earth cracks in Wupatki National
Mountain, Arizona, approximately 75
miles (121 kilometers (km)) south of
Grand Canyon National Park. Other
species in the genus Archeolarca have
been reported in California, Utah, Texas,
and Oregon (Muchmore 1981, p. 56;
Peck 1998, p. 23).
Distribution
The Cave of the Domes in Grand
Canyon National Park is currently the
only known location for the Grand
Canyon cave pseudoscorpion. Welbourn
(1978, pp. 36–41) conducted a regional
study of cave fauna on Horseshoe Mesa
of the Grand Canyon from 1977 to 1978.
Eight caves were examined including
Babylon Cave, Crystal Forest Cave,
Land’s End Cave, Middle Cave,
Scorpion Cave, Tse An Cho Cave,
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Tuning Fork Cave, and Cave of the
Domes. All caves except Land’s End
Cave and Scorpion Cave were visited
twice. On each visit, Welbourn (1978, p.
36) describes examining the walls,
ceilings, and floors for animals and
invertebrates. He identified 12
invertebrates from the 8 caves. The
Grand Canyon cave pseudoscorpion was
found only in the Cave of the Domes
(Welbourn 1978, pp. 38–41).
Wynne et al. (2008a, pp. 235–246)
summarized all published and
unpublished literature on cave-dwelling
invertebrates within Grand Canyon
National Park, as well as cave trip
reports on file at Grand Canyon National
Park Museum Collections. The literature
review examined 9 studies conducted
between 1975 and 2001 representing
surveys of 15 caves in Grand Canyon
National Park. Wynne et al. (2008a, pp.
237–238) reported 37 cave-dwelling
invertebrates with the Grand Canyon
cave pseudoscorpion referenced only in
the Cave of the Domes. This species may
possibly be endemic to Cave of the
Domes. In fact, a study of patterns of
endemism of eastern North American
cave fauna reported that within the
Pseudoscorpionida is a high level of
single-cave endemism compared to
other cave taxa (Christman et al. 2005,
pp. 1444, 1447). However, cave
biological research in Grand Canyon
National Park is quite limited (Wynne
2010, pers. comm.; Drost 2010, pers.
comm.) and more invertebrate surveys
need to be conducted before we can
conclude that the Grand Canyon cave
pseudoscorpion is endemic to the Cave
of the Domes. Further, because many
cave-dwelling organisms are hard to
find, cave inventories cannot be
considered complete without intensive
invertebrate trapping, baiting of the
entity, and multiple site visits (Wynne
2010, pers. comm.). We cannot describe
the distribution based on a single
specimen; therefore, we are not able to
determine the distribution of the Grand
Canyon cave pseudoscorpion.
Habitat and Biology
Most species of pseudoscorpions
occur in tropical and subtropical areas
throughout the world, although
pseudoscorpions can also be found in
temperate zones (Weygoldt 1969, pg.
108). They are found in a great variety
of habitats, but one essential feature
appears to be the presence of small
crevices where they can retreat. All
pseudoscorpion species spend most of
their lives within these crevices and
seldom appear on open ground. These
small crevices can be found in rocks,
tree bark, leaf litter, nests of birds and
other small mammals, and buildings.
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Another important habitat factor is
humidity; most pseudoscorpions prefer
high humidity, although some species
are found in arid conditions, such as
deserts (Weygoldt 1969, pp. 108–111).
There are few studies on the ecology
and habitat preferences of specific
species of pseudoscorpions. We have no
specific information about the habitat
and biology of the Grand Canyon cave
pseudoscorpion because the species is
known from only one specimen.
Accordingly, we can only speculate
about their habitat requirements and
biology based on the scant general
information known about
pseudoscorpions.
Welbourn (1978, p. 37) observed that
the single most important limiting factor
for the cave fauna on Horseshoe Mesa,
which includes Cave of the Domes, was
the lack of moisture. Welbourn (1978, p.
37) reported that most of the caves
surveyed were dry and dusty with low
relative humidity, and that most of the
caves examined, including Cave of the
Domes, received moisture from rainfall
that percolates through the limestone
above. Welbourn (1978, p. 40) reported
collecting the species ‘‘in the Cross
passage of Cave of the Domes in some
organic material (grass).’’ According to
the Grand Canyon National Park’s
hydrologist, the Cave of the Domes is
considered to be a dry cave with no
discharge or pools, but that the Cave of
the Domes has some small ephemeral
drip zones (Rice 2010, pers. comm.). We
do not know if the location where the
Grand Canyon cave pseudoscorpion was
found was optimal habitat or an
accidental find, but if pseudoscorpions
prefer humid locations, this location
may not represent optimal habitat.
Some species of pseudoscorpions are
known to be phoretic (use another
species for transportation) on other
arthropods such as flies, beetles, and
wasps. Pseudoscorpions will attach
themselves (not as parasites) to the legs
and appendages of the adult arthropod,
which permits them to ‘‘hitchhike.’’
According to Poinar et al. (1998, p. 79),
the principal benefit of pseudoscorpion
phoresy is dispersal; that is, to reach a
new habitat with an adequate supply of
food. We can only speculate on the
presence of the one specimen of Grand
Canyon cave pseudoscorpion in Cave of
the Domes, but perhaps it was carried
there and deposited by an arthropod.
The Park Service’s biological report
(Hill et al. 1998, pg. 16) from Cave of the
Domes indicated that packrat middens
(nests) were observed inside Cave of the
Domes. The report stated that a packrat
midden was found ‘‘in the second room
of the cave’’. Pseudoscorpions are often
present in rodent nests (Francke and
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´
Villegas-Guzman 2006, p. 289).
Muchmore (1991, pers. comm.) stated
that the genus Archeolarca does not
usually inhabit caves but rather is found
in packrat nests, although packrat
middens are sometimes found in caves.
´
Francke and Villegas-Guzman (2006, p.
297) conclude that pseudoscorpions
most likely coexist with a particular
rodent species in a mutualistic
association (a relationship between two
species where both species derive
benefits) in which pseudoscorpions feed
on adult and larval fleas, which reduces
the parasite load within the host nest.
The benefits to the pseudoscorpion
include the host nest providing suitable
microclimate, especially in semiarid
regions, as well as food (i.e., mites, fleas,
flies and their larvae). It is possible that
this species may be associated with
packrat middens or other small mammal
nests within Cave of the Domes, but we
cannot draw that conclusion based on
one specimen.
In summary, we lack sufficient
information on the species to reach
conclusions about the biology or the
habitat needs of the Grand Canyon cave
pseudoscorpion. This is primarily
because we know of only one specimen,
and we cannot make scientifically
sound conclusions regarding habitat
characteristics and biology based on a
single specimen.
Factors Affecting the Grand Canyon
Cave Pseudoscorpion
Section 4 of the Act (16 U.S.C. 1533)
and implementing regulations (50 CFR
part 424) set forth procedures for adding
species to, removing species from, or
reclassifying species on the Federal
Lists of Endangered and Threatened
Wildlife and Plants. Under section
4(a)(1) of the Act, a species may be
determined to be endangered or
threatened based on any of the
following five factors:
(A) The present or threatened
destruction, modification, or
curtailment of its habitat or range;
(B) Overutilization for commercial,
recreational, scientific, or educational
purposes;
(C) Disease or predation;
(D) The inadequacy of existing
regulatory mechanisms; or
(E) Other natural or manmade factors
affecting its continued existence.
In making this finding, information
pertaining to the Grand Canyon cave
pseudoscorpion in relation to the five
factors provided in section 4(a)(1) of the
Act is discussed below.
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Factor A. The Present or Threatened
Destruction, Modification, or
Curtailment of Its Habitat or Range
The Arizona Game and Fish
Department (2003, p. 2) stated that a
threat to the Grand Canyon cave
pseudoscorpion was groundwater
pollution. Our previous discussion in
the Habitat and Biology section
indicates that Cave of the Domes, where
the species was collected, is considered
dry and has very little ephemeral water
(i.e., small drip zones). Further, the
specimen was found associated with
dry, organic material (grass) in Cave of
the Domes. This description of the site
where the Grand Canyon cave
pseudoscorpion was collected seems
unlikely to be affected by groundwater
pollution because it is relatively dry,
and based on a single specimen we are
unable to determine the type of cave
habitat associated with this species.
Additionally, we have no specific
information regarding the presence or
introduction of contaminants or
pollutants in water sources on
Horseshoe Mesa, which could percolate
into the Cave of the Domes. Therefore,
we are unable to determine if
groundwater pollution is a threat.
Cave of the Domes is the only cave in
Grand Canyon National Park for which
visitation is allowed. It is unknown
whether recreation is modifying or
destroying the Grand Canyon cave
pseudoscorpion’s habitat. A report from
the Grand Canyon National Park
(Western Speleological Institute 1954,
pp. 1–2) stated that the cave has been
badly vandalized and floor deposits
have been marred by trampling.
However, it is unknown if this damage
affects the pseudoscorpion’s habitat
because that habitat is unknown. We
note that vandalism and trampling have
been identified as potential threats to
other pseudoscorpion species, such as
the Empire cave pseudoscorpion
(Microcreagris imperialis) (Muchmore
and Cokendolpher 1995, pp. 174–175)
and the Tooth cave pseudoscorpion
(Tartarocreagris texana) (Service 1994,
pp. 62–63). If the Grand Canyon cave
pseudoscorpion occupies packrat
middens, as do other pseudoscorpions
in the genus Archeolarca, then
recreational foot traffic may be a
discountable impact; however, we are
lacking sufficient information on the
habitat for this species. As such, we are
unable to determine if recreational
activity is affecting the Grand Canyon
cave pseudoscorpion or its habitat.
In summary, given the paucity of
biological information regarding the
Grand Canyon cave pseudoscorpion and
its habitat, we cannot determine or
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conclude that habitat degradation due to
groundwater pollution or recreational
activities in the Cave of the Domes is a
threat to the species now or in the
foreseeable future.
Factor B. Overutilization for
Commercial, Recreational, Scientific, or
Educational Purposes
Any commercial, recreational,
scientific, or educational collection
activities, including the collection of
pseudoscorpions, would require a
permit by the National Park Service.
Because of this regulation, there is no
data suggesting that overutilization for
commercial, recreational, scientific, or
educational purposes pose a threat to
the species. There are no known
commercial or recreational uses for
Grand Canyon cave pseudoscorpions.
Therefore, we find that the Grand
Canyon cave pseudoscorpion is not
threatened by overutilization now or in
the foreseeable future.
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Factor C. Disease or Predation
We have no information to indicate
that the Grand Canyon cave
pseudoscorpion is subject to disease or
predation. We have not encountered any
information that indicates the contrary;
however, in the absence of evidence that
this factor may constitute a threat to the
species, we cannot determine or
conclude that the Grand Canyon cave
pseudoscorpion is threatened by disease
or predation now or in the foreseeable
future.
Factor D. The Inadequacy of Existing
Regulatory Mechanisms
Under the current National Park
Service policy, all caves in Grand
Canyon National Park are closed to
visitation by recreational users except
for the Cave of the Domes. The Park
Service has the authority, under the
Federal Cave Resources Protection Act
of 1988, to close areas to visitors if there
is significant degradation of a resource
or the threat of degradation or damage.
On Park Service lands, all caves are
deemed ‘‘significant,’’ and the Park
Service protects the caves, including
biological, cultural, and paleontological
resources within the caves. The decision
to regulate visitors or close the cave to
recreational use is made by the Park
Superintendent with supporting
documentation from resource managers.
Official criteria for determining
recreational access to Grand Canyon
National Park caves has not been
established, but the initiation of a Cave
Management Plan is planned.
Cave of the Domes is located beneath
Horseshoe Mesa and is well known to
hikers and cavers. The Web site
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https://www.birdandhike.com provides a
detailed overview of the cave, including
photos and directions to the trailhead
and to the mouth of the cave. The Web
site https://www.kaibab.org also provides
information about Cave of the Domes
and states that many formations have
been damaged by careless individuals
and asks visitors to treat the cave with
respect. As stated above, we lack data to
assess the effect of recreation on the
petitioned species. Therefore, due to the
lack of information regarding impacts of
recreational visitors and the Park
Service’s ability to close the area if
additional information comes to light,
we find that the Grand Canyon cave
pseudoscorpion is not threatened by
inadequacy of regulatory mechanisms
now or in the foreseeable future.
Factor E. Other Natural or Manmade
Factors Affecting Its Continued
Existence
Model predictions are that
temperatures in the Southwestern
United States will continue to increase,
with extreme weather events (such as
heat waves, drought, and flooding)
occurring with more frequency due to
global climate change (Archer and
Predick 2008, p. 24). It is unknown how
cave-adapted taxa will respond to global
climate change. Baseline information on
ecosystem structure and community
structure is lacking for many caves, and
we do not know how cave-dwelling
species will respond to rising
temperatures. Different layers of a cave
may be affected differently, depending
on their depth. (Wynne et al. 2008b, p.
241). There will most likely be a lag
effect; caves with shallow vertical depth
are predicted to have a more immediate
response than caves with deeper vertical
depth (Wynne 2010, pers. comm.). We
have no information on the geophysical
properties of Cave of the Domes.
Researchers are currently attempting to
understand the geophysical properties
of caves as they relate to cave depth, the
potential effects of rising surface
temperatures on cave temperatures, and
how the physiological requirements of
cave-dwelling and cave-adapted species
are affected by climate change (Drost
2010, pers. comm.). Based on the best
available information, we cannot
determine or conclude that climate
change is a threat to the Grand Canyon
cave pseudoscorpion now or in the
foreseeable future.
Finding
As required by the Act, we considered
the five factors in assessing whether the
Grand Canyon cave pseudoscorpion is
threatened or endangered throughout all
or a significant portion of its range. We
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examined the best scientific and
commercial information available
regarding the past, present, and future
threats faced by the Grand Canyon cave
pseudoscorpion. We reviewed the
petition, information available in our
files, other available published and
unpublished information, and we
consulted with recognized invertebrate
experts and the Grand Canyon National
Park biologist and hydrologist.
In considering what factors might
constitute threats, we must look beyond
the mere exposure of the species to the
factor to determine whether the species
responds to the factor in a way that
causes actual impacts to the species. If
there is exposure to a factor, but no
response, or only a positive response,
that factor is not a threat. If there is
exposure and the species responds
negatively, the factor may be a threat
and we then attempt to determine how
significant a threat it is. If the threat is
significant, it may drive or contribute to
the risk of extinction of the species such
that the species warrants listing as
threatened or endangered as those terms
are defined by the Act. This does not
necessarily require empirical proof of a
threat. The combination of exposure and
some corroborating evidence of how the
species is likely impacted could suffice.
The mere identification of factors that
could impact a species negatively is not
sufficient to compel a finding that
listing is appropriate; we require
evidence that these factors are operative
threats that act on the species to the
point that the species meets the
definition of threatened or endangered
under the Act.
Based on our review of the best
available scientific and commercial
information pertaining to the five
factors, we found no evidence to
indicate that there are threats to the
species or its habitat, from any of the
five factors. For this reason, we
conclude that the Grand Canyon cave
pseudoscorpion does not meet the
definition of a threatened or endangered
species and are, therefore,
recommending a finding of ‘‘not
warranted.’’
We request that you submit any new
information concerning the distribution
and status of, or threats to, the Grand
Canyon cave pseudoscorpion to our U.S.
Fish and Wildlife Service Office (see
ADDRESSES) whenever it becomes
available. New information will help us
monitor the Grand Canyon cave
pseudoscorpion and encourage its
conservation. If an emergency situation
develops for the Grand Canyon cave
pseudoscorpion or any other species, we
will act to provide immediate
protection.
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42658
Federal Register / Vol. 76, No. 138 / Tuesday, July 19, 2011 / Proposed Rules
References Cited
A complete list of references cited is
available on the Internet at https://
www.regulations.gov and upon request
from the U.S. Fish and Wildlife Service,
Arizona Ecological Services Field Office
(see ADDRESSES).
Author(s)
The primary authors of this finding
are the staff members of the Arizona
Ecological Services Field Office.
Authority: The authority for this section is
section 4 of the Endangered Species Act of
1973, as amended (16 U.S.C. 1531 et seq.).
Dated: June 21, 2011.
Rowan W. Gould,
Acting Director, Fish and Wildlife Service.
[FR Doc. 2011–17864 Filed 7–18–11; 8:45 am]
BILLING CODE 4310–55–P
DEPARTMENT OF COMMERCE
National Oceanic and Atmospheric
Administration
50 CFR Part 223
[Docket No. 110615334–1325–01]
RIN 0648–XA311
Endangered and Threatened Species:
Authorizing Release of a Nonessential
Experimental Population of Upper
Columbia Spring-Run Chinook Salmon
in the Okanogan River Basin Under the
Endangered Species Act
srobinson on DSK4SPTVN1PROD with PROPOSALS
AGENCY: National Marine Fisheries
Service (NMFS), National Oceanic and
Atmospheric Administration (NOAA),
Commerce.
ACTION: Advance Notice of Proposed
Rulemaking; Request for Information.
SUMMARY: We, the National Marine
Fisheries Service (NMFS), will be
considering a proposal to authorize a
nonessential experimental population of
Upper Columbia (UC) spring-run
Chinook salmon (Oncorhynchus
tshawytscha) in the Okanogan River and
its tributaries in Okanogan County,
Washington under the Endangered
Species Act (ESA) of 1973, as amended.
The geographic boundaries of the
experimental population area would
likely include the entire Okanogan River
subbasin and a portion of the mainstem
Columbia River from the confluence of
the Columbia and Okanogan Rivers
upstream to the base of Chief Joseph
Dam. We will consider the best
available information to determine if
reintroduction of Chinook salmon is
biologically feasible and will promote
the conservation of the UC spring-run
VerDate Mar<15>2010
17:35 Jul 18, 2011
Jkt 223001
Chinook salmon Evolutionarily
Significant Unit (ESU). This advance
notice of proposed rulemaking (ANPR)
identifies policy and technical issues for
consideration and evaluation, and
solicits comments regarding them.
DATES: Comments and information
regarding the designation process may
be sent to us (see ADDRESSES), no later
than 5 p.m. Pacific Time on September
19, 2011.
ADDRESSES: Comments may be sent to
Chief, Protected Resources Division,
NMFS, 1201 NE. Lloyd Blvd.—Suite
1100, Portland, OR 97232. Comments
may also be sent via facsimile (fax) to
503–230–5441 or submitted on the
Internet via the Federal Rulemaking
portal at https://www.regulations.gov.
Follow the instructions for submitting
comments.
Instructions: All comments received
are a part of the public record and will
generally be posted to https://
www.regulations.gov without change.
We may elect not to post comments that
contain obscene or threatening content.
All personal identifying information (for
example, name, address, etc.)
voluntarily submitted by the commenter
may be publicly accessible. Do not
submit Confidential Business
Information or otherwise sensitive or
protected information.
We will accept anonymous comments
(enter N/A in the required fields, if you
wish to remain anonymous). You may
submit attachments to electronic
comments in Microsoft Word, Excel,
WordPerfect, or Adobe PDF file formats
only.
FOR FURTHER INFORMATION CONTACT: Eric
Murray, NMFS, Northwest Region,
Portland, OR 503–231–2378; or Dwayne
Meadows, NMFS, Office of Protected
Resources, Silver Spring, MD 301–713–
1401.
SUPPLEMENTARY INFORMATION:
Rulemaking Background
We first listed the Upper Columbia
(UC) spring-run Chinook salmon ESU as
endangered under the ESA on March 24,
1999 (64 FR 14308), and reaffirmed this
status on June 28, 2005 (70 FR 37160).
ESA Section 9 ‘‘take’’ prohibitions
currently apply to the UC spring-run
Chinook salmon ESU because of its
endangered status.
The listed ESU currently includes all
naturally spawned populations of
spring-run Chinook salmon in
accessible reaches of Columbia River
tributaries between Rock Island and
Chief Joseph Dams, excluding the
Okanogan River. Listed spring-run
Chinook salmon from this ESU
currently spawn in three river basins in
PO 00000
Frm 00069
Fmt 4702
Sfmt 4702
eastern Washington: The Methow,
Entiat and Wenatchee. A fourth
population historically inhabited the
Okanogan River Basin, but was
extirpated in the 1930s because of
overfishing, hydropower development,
and habitat degradation (NMFS, 2007).
The designated critical habitat of UC
spring-run Chinook salmon similarly
includes all accessible reaches of
Columbia River tributaries between
Rock Island and Chief Joseph Dams, but
excludes the Okanogan River. We did
not include the Okanogan River Basin in
any critical habitat designation because
the Okanogan population of spring-run
Chinook salmon no longer existed.
The listed UC spring-run Chinook
salmon ESU also includes six artificial
propagation programs: The Twisp River,
Chewuch River, Methow Composite,
Winthrop National Fish Hatchery,
Chiwawa River, and White River spring
Chinook salmon hatchery programs.
On October 9, 2007, we adopted a
final recovery plan for the UC springrun Chinook salmon ESU (72 FR 57303).
The recovery plan identifies three extant
populations in this ESU (the Methow,
Wenatchee, and Entiat) and an historic,
extirpated population in the Okanogan
River Basin (NMFS, 2007). The recovery
plan identifies re-establishment of a
population in the Okanogan River Basin
as a recovery action (NMFS, 2007). Reestablishment of a spring-run Chinook
salmon population in the Okanogan
River Basin could aid recovery of this
ESU by increasing abundance, by
improving spatial structure, and by
reducing the risk of extinction to the
ESU as a whole.
On November 22, 2010, we received
a letter from the Confederated Tribes of
the Colville Reservation (CTCR)
requesting that we authorize the release
of an experimental population of springrun Chinook salmon in the Okanogan
River Basin. The CTCR has also initiated
discussions on this topic with the U.S.
Fish and Wildlife Service (USFWS), the
Bonneville Power Administration, the
Army Corps of Engineers, the Bureau of
Reclamation, the Washington
Department of Fish and Wildlife
(WDFW), and the Okanagan Nations
Alliance of Canada. The CTCR’s request
included a large amount of information
on the biology of UC spring-run
Chinook salmon and the possible
management implications of releasing
an experimental population in the
Okanogan Basin.
Statutory and Regulatory Framework
Section 10(j) of the ESA allows the
Secretary of Commerce (Secretary) to
authorize the release of populations of
listed species outside their current range
E:\FR\FM\19JYP1.SGM
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Agencies
[Federal Register Volume 76, Number 138 (Tuesday, July 19, 2011)]
[Proposed Rules]
[Pages 42654-42658]
From the Federal Register Online via the Government Printing Office [www.gpo.gov]
[FR Doc No: 2011-17864]
-----------------------------------------------------------------------
DEPARTMENT OF THE INTERIOR
Fish and Wildlife Service
50 CFR Part 17
[Docket No. FWS-R2-ES-2011-0044; MO 92210-0-0008-B2]
Endangered and Threatened Wildlife and Plants; Petition To List
Grand Canyon Cave Pseudoscorpion
AGENCY: Fish and Wildlife Service, Interior.
ACTION: Notice of 12-month petition finding.
-----------------------------------------------------------------------
SUMMARY: We, the U.S. Fish and Wildlife Service (Service), announce a
12-month finding on a petition to list the Grand Canyon cave
pseudoscorpion (Archeolarca cavicola) as threatened or endangered with
critical habitat under the Endangered Species Act of 1973, as amended
(Act). After review of the best scientific and commercial information
available, we find that listing the Grand Canyon cave pseudoscorpion is
not warranted at this time. However, we ask the public to submit to us
any new information that becomes available concerning the threats to
the Grand Canyon cave pseudoscorpion or its habitat at any time.
DATES: The finding announced in this document was made on July 19,
2011.
ADDRESSES: This finding is available on the Internet at https://www.regulations.gov at Docket Number FWS-R2-ES-2011-0044. Supporting
documentation we used in preparing this finding is available for public
inspection, by appointment, during normal business hours by contacting
the U.S. Fish and Wildlife Service, Arizona Ecological Services Field
Office, 2321 W. Royal Palm Road, Suite 103, Phoenix, AZ 85021;
telephone (602) 242-0210; facsimile (602) 242-2513. If you use a
telecommunications device for the deaf (TDD), please call the Federal
Information Relay Service (FIRS) at (800) 877-8339. Please submit any
new information, comments, or questions concerning this finding to the
above address.
FOR FURTHER INFORMATION CONTACT: Steve Spangle, Field Supervisor, U.S.
Fish and Wildlife Service, Arizona Ecological Services Field Office,
2321 W. Royal Palm Road, Suite 103, Phoenix, AZ 85021; telephone (602)
242-0210; facsimile (602) 242-2513. If you use a telecommunications
device for the deaf (TDD), please call the Federal Information Relay
Service (FIRS) at (800) 877-8339.
SUPPLEMENTARY INFORMATION:
Background
Section 4(b)(3)(B) of the Endangered Species Act of 1973, as
amended (Act) (16 U.S.C. 1531 et seq.), requires that, for any petition
containing substantial scientific or commercial information indicating
that listing the species may be warranted, we make a finding within 12
months of the date of receipt of the petition. In this finding we
determine that the petitioned action is: (a) Not warranted, (b)
warranted, or (c) warranted, but immediate proposal of a regulation
implementing the petitioned action is precluded by other pending
proposals to determine whether species are threatened or endangered,
and expeditious progress is being made to add or remove qualified
species from the Lists of Endangered and Threatened Wildlife and
Plants. Section 4(b)(3)(C) of the Act requires that we treat a petition
for which the requested action is found to be warranted but precluded
as though resubmitted on the date of such finding, that is, requiring a
subsequent finding to be made within 12 months. We must publish these
12-month findings in the Federal Register.
Previous Federal Actions
The Grand Canyon cave pseudoscorpion was formerly a candidate 2
species, a taxon for which information in our possession indicated that
proposing to list was possibly appropriate, but for which persuasive
data on biological vulnerability and threats were not available to
support a proposed listing rule (54 FR 554; January 6, 1989). The
designation of candidate 2 species was discontinued in
[[Page 42655]]
1996; therefore, the species has no candidate status.
On June 25, 2007, we received a formal petition dated June 18,
2007, from Forest Guardians (now WildEarth Guardians), requesting that
we do the following: (1) Consider all full species in our Southwest
Region ranked as G1 or G1G2 by the organization NatureServe for
listing, except those that are currently listed, proposed for listing,
or candidates; and (2) list each species under the Act as either
endangered or threatened with critical habitat. The petitioners
presented two tables that collectively listed 475 species for
consideration and requested that the Service incorporate all analyses,
references, and documentation provided by NatureServe in its online
database https://www.natureserve.org/ into the petition. The petition
clearly identified itself as a petition and included the appropriate
identification information, as required in 50 CFR 424.14(a). We
acknowledged the receipt of the petition in a letter to WildEarth
Guardians dated July 11, 2007.
On December 16, 2009, we made a 90-day finding that the petition
presented substantial scientific information indicating that listing 67
of the 475 species may be warranted; the Grand Canyon cave
pseudoscorpion (incorrectly referenced as the Grand Canyon cave
scorpion) was in that group of 67 species. Based on the evaluation of
the information provided in the petition, we determined that the
petition presented substantial information to indicate that listing the
Grand Canyon cave pseudoscorpion may be warranted due to the present or
threatened destruction, modification, or curtailment of its habitat or
range resulting from groundwater pollution and recreational impacts,
and to the inadequacy of existing regulatory mechanisms resulting from
unregulated visitation. The 90-day finding and our initiation of a
status review was published in the Federal Register on December 16,
2009 (74 FR 66866). This notice constitutes the 12-month finding on the
June 18, 2007, petition to list the Grand Canyon cave pseudoscorpion as
threatened or endangered.
Species Information
Species Description
In 1978 W. Calvin Welbourn collected one female specimen of the
Grand Canyon cave pseudoscorpion (Class Arachnida, Order
Pseudoscorpionida, Family Garypidae) in Grand Canyon National Park's
Cave of the Domes. This specimen was first described by Muchmore (1981,
p. 55). Welbourn's unpublished report (1978, p. 40) stated that the
specimen was an undescribed troglophile. A troglophile is a species
that can spend its entire life within caves, does not exhibit
adaptations for living in caves, but can also be found in suitable
habitats outside of caves, such as mines or animal burrows. This is in
comparison to troglobites, which are species that are found exclusively
in caves and have developed adaptations for cave life, such as
heightened sense of hearing, touch, and smell. The Grand Canyon cave
pseudoscorpion was recognized within the genus Archeolarca and was
given the species name cavicola in recognition of its subterranean
habitat where it was collected by Welbourn (1981, p. 55). The specimen
collected in 1978 is the only one known to exist. No other individuals
are known to have been collected since 1978, although very little
effort has been made to collect this or other species in the genus
(Service 1991, p. 3).
Pseudoscorpions are tiny arachnids bearing large chelae, or claws,
but lacking a telson, or stinger, that true scorpions possess. The
specimen of Grand Canyon cave pseudoscorpion, when compared to other
species of pseudoscorpions, such as Archeolarca welbourni and
Archeolarca guadalupensis, was considered large (0.12 inches (in) or
3.03 millimeters (mm)), had longer appendages, more reduced posterior
eyes, and fewer setae (stiff bristles present on the body) on its upper
dorsal section (Muchmore 1981, p. 56). Muchmore (1981, pp. 52-56)
described three new species of Archeolarca, and concluded that the
Grand Canyon cave pseudoscorpion showed the greatest overall adaption
to the cave environment. Welbourn (1978, p. 40) noted that the specimen
appeared to be similar to the pseudoscorpions from earth cracks in
Wupatki National Mountain, Arizona, approximately 75 miles (121
kilometers (km)) south of Grand Canyon National Park. Other species in
the genus Archeolarca have been reported in California, Utah, Texas,
and Oregon (Muchmore 1981, p. 56; Peck 1998, p. 23).
Distribution
The Cave of the Domes in Grand Canyon National Park is currently
the only known location for the Grand Canyon cave pseudoscorpion.
Welbourn (1978, pp. 36-41) conducted a regional study of cave fauna on
Horseshoe Mesa of the Grand Canyon from 1977 to 1978. Eight caves were
examined including Babylon Cave, Crystal Forest Cave, Land's End Cave,
Middle Cave, Scorpion Cave, Tse An Cho Cave, Tuning Fork Cave, and Cave
of the Domes. All caves except Land's End Cave and Scorpion Cave were
visited twice. On each visit, Welbourn (1978, p. 36) describes
examining the walls, ceilings, and floors for animals and
invertebrates. He identified 12 invertebrates from the 8 caves. The
Grand Canyon cave pseudoscorpion was found only in the Cave of the
Domes (Welbourn 1978, pp. 38-41).
Wynne et al. (2008a, pp. 235-246) summarized all published and
unpublished literature on cave-dwelling invertebrates within Grand
Canyon National Park, as well as cave trip reports on file at Grand
Canyon National Park Museum Collections. The literature review examined
9 studies conducted between 1975 and 2001 representing surveys of 15
caves in Grand Canyon National Park. Wynne et al. (2008a, pp. 237-238)
reported 37 cave-dwelling invertebrates with the Grand Canyon cave
pseudoscorpion referenced only in the Cave of the Domes. This species
may possibly be endemic to Cave of the Domes. In fact, a study of
patterns of endemism of eastern North American cave fauna reported that
within the Pseudoscorpionida is a high level of single-cave endemism
compared to other cave taxa (Christman et al. 2005, pp. 1444, 1447).
However, cave biological research in Grand Canyon National Park is
quite limited (Wynne 2010, pers. comm.; Drost 2010, pers. comm.) and
more invertebrate surveys need to be conducted before we can conclude
that the Grand Canyon cave pseudoscorpion is endemic to the Cave of the
Domes. Further, because many cave-dwelling organisms are hard to find,
cave inventories cannot be considered complete without intensive
invertebrate trapping, baiting of the entity, and multiple site visits
(Wynne 2010, pers. comm.). We cannot describe the distribution based on
a single specimen; therefore, we are not able to determine the
distribution of the Grand Canyon cave pseudoscorpion.
Habitat and Biology
Most species of pseudoscorpions occur in tropical and subtropical
areas throughout the world, although pseudoscorpions can also be found
in temperate zones (Weygoldt 1969, pg. 108). They are found in a great
variety of habitats, but one essential feature appears to be the
presence of small crevices where they can retreat. All pseudoscorpion
species spend most of their lives within these crevices and seldom
appear on open ground. These small crevices can be found in rocks, tree
bark, leaf litter, nests of birds and other small mammals, and
buildings.
[[Page 42656]]
Another important habitat factor is humidity; most pseudoscorpions
prefer high humidity, although some species are found in arid
conditions, such as deserts (Weygoldt 1969, pp. 108-111).
There are few studies on the ecology and habitat preferences of
specific species of pseudoscorpions. We have no specific information
about the habitat and biology of the Grand Canyon cave pseudoscorpion
because the species is known from only one specimen. Accordingly, we
can only speculate about their habitat requirements and biology based
on the scant general information known about pseudoscorpions.
Welbourn (1978, p. 37) observed that the single most important
limiting factor for the cave fauna on Horseshoe Mesa, which includes
Cave of the Domes, was the lack of moisture. Welbourn (1978, p. 37)
reported that most of the caves surveyed were dry and dusty with low
relative humidity, and that most of the caves examined, including Cave
of the Domes, received moisture from rainfall that percolates through
the limestone above. Welbourn (1978, p. 40) reported collecting the
species ``in the Cross passage of Cave of the Domes in some organic
material (grass).'' According to the Grand Canyon National Park's
hydrologist, the Cave of the Domes is considered to be a dry cave with
no discharge or pools, but that the Cave of the Domes has some small
ephemeral drip zones (Rice 2010, pers. comm.). We do not know if the
location where the Grand Canyon cave pseudoscorpion was found was
optimal habitat or an accidental find, but if pseudoscorpions prefer
humid locations, this location may not represent optimal habitat.
Some species of pseudoscorpions are known to be phoretic (use
another species for transportation) on other arthropods such as flies,
beetles, and wasps. Pseudoscorpions will attach themselves (not as
parasites) to the legs and appendages of the adult arthropod, which
permits them to ``hitchhike.'' According to Poinar et al. (1998, p.
79), the principal benefit of pseudoscorpion phoresy is dispersal; that
is, to reach a new habitat with an adequate supply of food. We can only
speculate on the presence of the one specimen of Grand Canyon cave
pseudoscorpion in Cave of the Domes, but perhaps it was carried there
and deposited by an arthropod.
The Park Service's biological report (Hill et al. 1998, pg. 16)
from Cave of the Domes indicated that packrat middens (nests) were
observed inside Cave of the Domes. The report stated that a packrat
midden was found ``in the second room of the cave''. Pseudoscorpions
are often present in rodent nests (Francke and Villegas-Guzm[aacute]n
2006, p. 289). Muchmore (1991, pers. comm.) stated that the genus
Archeolarca does not usually inhabit caves but rather is found in
packrat nests, although packrat middens are sometimes found in caves.
Francke and Villegas-Guzm[aacute]n (2006, p. 297) conclude that
pseudoscorpions most likely coexist with a particular rodent species in
a mutualistic association (a relationship between two species where
both species derive benefits) in which pseudoscorpions feed on adult
and larval fleas, which reduces the parasite load within the host nest.
The benefits to the pseudoscorpion include the host nest providing
suitable microclimate, especially in semiarid regions, as well as food
(i.e., mites, fleas, flies and their larvae). It is possible that this
species may be associated with packrat middens or other small mammal
nests within Cave of the Domes, but we cannot draw that conclusion
based on one specimen.
In summary, we lack sufficient information on the species to reach
conclusions about the biology or the habitat needs of the Grand Canyon
cave pseudoscorpion. This is primarily because we know of only one
specimen, and we cannot make scientifically sound conclusions regarding
habitat characteristics and biology based on a single specimen.
Factors Affecting the Grand Canyon Cave Pseudoscorpion
Section 4 of the Act (16 U.S.C. 1533) and implementing regulations
(50 CFR part 424) set forth procedures for adding species to, removing
species from, or reclassifying species on the Federal Lists of
Endangered and Threatened Wildlife and Plants. Under section 4(a)(1) of
the Act, a species may be determined to be endangered or threatened
based on any of the following five factors:
(A) The present or threatened destruction, modification, or
curtailment of its habitat or range;
(B) Overutilization for commercial, recreational, scientific, or
educational purposes;
(C) Disease or predation;
(D) The inadequacy of existing regulatory mechanisms; or
(E) Other natural or manmade factors affecting its continued
existence.
In making this finding, information pertaining to the Grand Canyon
cave pseudoscorpion in relation to the five factors provided in section
4(a)(1) of the Act is discussed below.
Factor A. The Present or Threatened Destruction, Modification, or
Curtailment of Its Habitat or Range
The Arizona Game and Fish Department (2003, p. 2) stated that a
threat to the Grand Canyon cave pseudoscorpion was groundwater
pollution. Our previous discussion in the Habitat and Biology section
indicates that Cave of the Domes, where the species was collected, is
considered dry and has very little ephemeral water (i.e., small drip
zones). Further, the specimen was found associated with dry, organic
material (grass) in Cave of the Domes. This description of the site
where the Grand Canyon cave pseudoscorpion was collected seems unlikely
to be affected by groundwater pollution because it is relatively dry,
and based on a single specimen we are unable to determine the type of
cave habitat associated with this species. Additionally, we have no
specific information regarding the presence or introduction of
contaminants or pollutants in water sources on Horseshoe Mesa, which
could percolate into the Cave of the Domes. Therefore, we are unable to
determine if groundwater pollution is a threat.
Cave of the Domes is the only cave in Grand Canyon National Park
for which visitation is allowed. It is unknown whether recreation is
modifying or destroying the Grand Canyon cave pseudoscorpion's habitat.
A report from the Grand Canyon National Park (Western Speleological
Institute 1954, pp. 1-2) stated that the cave has been badly vandalized
and floor deposits have been marred by trampling. However, it is
unknown if this damage affects the pseudoscorpion's habitat because
that habitat is unknown. We note that vandalism and trampling have been
identified as potential threats to other pseudoscorpion species, such
as the Empire cave pseudoscorpion (Microcreagris imperialis) (Muchmore
and Cokendolpher 1995, pp. 174-175) and the Tooth cave pseudoscorpion
(Tartarocreagris texana) (Service 1994, pp. 62-63). If the Grand Canyon
cave pseudoscorpion occupies packrat middens, as do other
pseudoscorpions in the genus Archeolarca, then recreational foot
traffic may be a discountable impact; however, we are lacking
sufficient information on the habitat for this species. As such, we are
unable to determine if recreational activity is affecting the Grand
Canyon cave pseudoscorpion or its habitat.
In summary, given the paucity of biological information regarding
the Grand Canyon cave pseudoscorpion and its habitat, we cannot
determine or
[[Page 42657]]
conclude that habitat degradation due to groundwater pollution or
recreational activities in the Cave of the Domes is a threat to the
species now or in the foreseeable future.
Factor B. Overutilization for Commercial, Recreational, Scientific, or
Educational Purposes
Any commercial, recreational, scientific, or educational collection
activities, including the collection of pseudoscorpions, would require
a permit by the National Park Service. Because of this regulation,
there is no data suggesting that overutilization for commercial,
recreational, scientific, or educational purposes pose a threat to the
species. There are no known commercial or recreational uses for Grand
Canyon cave pseudoscorpions. Therefore, we find that the Grand Canyon
cave pseudoscorpion is not threatened by overutilization now or in the
foreseeable future.
Factor C. Disease or Predation
We have no information to indicate that the Grand Canyon cave
pseudoscorpion is subject to disease or predation. We have not
encountered any information that indicates the contrary; however, in
the absence of evidence that this factor may constitute a threat to the
species, we cannot determine or conclude that the Grand Canyon cave
pseudoscorpion is threatened by disease or predation now or in the
foreseeable future.
Factor D. The Inadequacy of Existing Regulatory Mechanisms
Under the current National Park Service policy, all caves in Grand
Canyon National Park are closed to visitation by recreational users
except for the Cave of the Domes. The Park Service has the authority,
under the Federal Cave Resources Protection Act of 1988, to close areas
to visitors if there is significant degradation of a resource or the
threat of degradation or damage. On Park Service lands, all caves are
deemed ``significant,'' and the Park Service protects the caves,
including biological, cultural, and paleontological resources within
the caves. The decision to regulate visitors or close the cave to
recreational use is made by the Park Superintendent with supporting
documentation from resource managers. Official criteria for determining
recreational access to Grand Canyon National Park caves has not been
established, but the initiation of a Cave Management Plan is planned.
Cave of the Domes is located beneath Horseshoe Mesa and is well
known to hikers and cavers. The Web site https://www.birdandhike.com
provides a detailed overview of the cave, including photos and
directions to the trailhead and to the mouth of the cave. The Web site
https://www.kaibab.org also provides information about Cave of the Domes
and states that many formations have been damaged by careless
individuals and asks visitors to treat the cave with respect. As stated
above, we lack data to assess the effect of recreation on the
petitioned species. Therefore, due to the lack of information regarding
impacts of recreational visitors and the Park Service's ability to
close the area if additional information comes to light, we find that
the Grand Canyon cave pseudoscorpion is not threatened by inadequacy of
regulatory mechanisms now or in the foreseeable future.
Factor E. Other Natural or Manmade Factors Affecting Its Continued
Existence
Model predictions are that temperatures in the Southwestern United
States will continue to increase, with extreme weather events (such as
heat waves, drought, and flooding) occurring with more frequency due to
global climate change (Archer and Predick 2008, p. 24). It is unknown
how cave-adapted taxa will respond to global climate change. Baseline
information on ecosystem structure and community structure is lacking
for many caves, and we do not know how cave-dwelling species will
respond to rising temperatures. Different layers of a cave may be
affected differently, depending on their depth. (Wynne et al. 2008b, p.
241). There will most likely be a lag effect; caves with shallow
vertical depth are predicted to have a more immediate response than
caves with deeper vertical depth (Wynne 2010, pers. comm.). We have no
information on the geophysical properties of Cave of the Domes.
Researchers are currently attempting to understand the geophysical
properties of caves as they relate to cave depth, the potential effects
of rising surface temperatures on cave temperatures, and how the
physiological requirements of cave-dwelling and cave-adapted species
are affected by climate change (Drost 2010, pers. comm.). Based on the
best available information, we cannot determine or conclude that
climate change is a threat to the Grand Canyon cave pseudoscorpion now
or in the foreseeable future.
Finding
As required by the Act, we considered the five factors in assessing
whether the Grand Canyon cave pseudoscorpion is threatened or
endangered throughout all or a significant portion of its range. We
examined the best scientific and commercial information available
regarding the past, present, and future threats faced by the Grand
Canyon cave pseudoscorpion. We reviewed the petition, information
available in our files, other available published and unpublished
information, and we consulted with recognized invertebrate experts and
the Grand Canyon National Park biologist and hydrologist.
In considering what factors might constitute threats, we must look
beyond the mere exposure of the species to the factor to determine
whether the species responds to the factor in a way that causes actual
impacts to the species. If there is exposure to a factor, but no
response, or only a positive response, that factor is not a threat. If
there is exposure and the species responds negatively, the factor may
be a threat and we then attempt to determine how significant a threat
it is. If the threat is significant, it may drive or contribute to the
risk of extinction of the species such that the species warrants
listing as threatened or endangered as those terms are defined by the
Act. This does not necessarily require empirical proof of a threat. The
combination of exposure and some corroborating evidence of how the
species is likely impacted could suffice. The mere identification of
factors that could impact a species negatively is not sufficient to
compel a finding that listing is appropriate; we require evidence that
these factors are operative threats that act on the species to the
point that the species meets the definition of threatened or endangered
under the Act.
Based on our review of the best available scientific and commercial
information pertaining to the five factors, we found no evidence to
indicate that there are threats to the species or its habitat, from any
of the five factors. For this reason, we conclude that the Grand Canyon
cave pseudoscorpion does not meet the definition of a threatened or
endangered species and are, therefore, recommending a finding of ``not
warranted.''
We request that you submit any new information concerning the
distribution and status of, or threats to, the Grand Canyon cave
pseudoscorpion to our U.S. Fish and Wildlife Service Office (see
ADDRESSES) whenever it becomes available. New information will help us
monitor the Grand Canyon cave pseudoscorpion and encourage its
conservation. If an emergency situation develops for the Grand Canyon
cave pseudoscorpion or any other species, we will act to provide
immediate protection.
[[Page 42658]]
References Cited
A complete list of references cited is available on the Internet at
https://www.regulations.gov and upon request from the U.S. Fish and
Wildlife Service, Arizona Ecological Services Field Office (see
ADDRESSES).
Author(s)
The primary authors of this finding are the staff members of the
Arizona Ecological Services Field Office.
Authority: The authority for this section is section 4 of the
Endangered Species Act of 1973, as amended (16 U.S.C. 1531 et seq.).
Dated: June 21, 2011.
Rowan W. Gould,
Acting Director, Fish and Wildlife Service.
[FR Doc. 2011-17864 Filed 7-18-11; 8:45 am]
BILLING CODE 4310-55-P